Castilleja subinclusa |
Castilleja chrymactis |
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Franciscan paint brush, longleaf Indian paintbrush |
green Indian paintbrush, green paintbrush |
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Habit | Herbs, perennial, 1.8–12 dm; from a woody caudex; with a taproot. | Herbs, perennial, 2.8–6.6 dm; from a woody caudex; with a taproot. | ||||||||
Stems | solitary or few to many, erect to ascending, often leaning on nearby shrubs (var. jepsonii), unbranched or branched, often with small, leafy axillary shoots, hairs matted to spreading, short and long, soft, mixed with short stipitate-glandular ones. |
several to many, erect to ascending, unbranched or branched, sometimes with short, leafy axillary shoots, glabrous or hairs sparse proximally, distally spreading-erect, long, soft, eglandular, mixed with shorter ones. |
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Leaves | green, pale gray-green, sometimes dull reddish purple, linear to narrowly, sometimes broadly, lanceolate, 2–10.5 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0(–3)-lobed, apex acuminate to acute; lobes spreading, linear to short-lanceolate, often arising near mid length, apex acute to obtuse. |
green or purple-tinged, sometimes deep purple, linear-lanceolate to lanceolate or ovate, (1.5–)5–11.7 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0–3(–5)-lobed, apex acute to obtuse, sometimes rounded; lobes erect or ascending, linear, apex acute or acuminate. |
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Inflorescences | 3–30 × 1–7 cm; bracts green or dull purple throughout, or proximally green or dull purple, distally bright red to orange-red, sometimes salmon, orange, or yellow, usually linear to lanceolate, distal sometimes broadly lanceolate to oblong or elliptic, 0(–3)-lobed; lobes spreading to erect, linear to oblong or triangular, short or medium length, arising near or above mid length, apex acute to obtuse. |
3.5–18 × 3–7.5 cm; bracts proximally pale green to dull purplish, distally bright red or scarlet to yellow, orange, yellow-orange, and salmon, sometimes tricolored with central band of deeper purple or deeper shade of the far-distal color, broadly lanceolate, oblong, elliptic, or oblong, 0–3(–9)-lobed; lobes ascending, lanceolate, triangular, or oblong, short, arising at or above mid length, central lobe apex rounded, lateral ones acute to obtuse. |
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Corollas | curved proximally, straight to slightly curved distally, 25–57 mm; tube 15–29 mm; beak subequal to or exceeding calyx but abaxial lip, beak, and often part of tube usually exserted out abaxial cleft; beak adaxially green, yellow-green, or yellow, 13–21 mm, margins red, orange, or yellow; abaxial lip green, yellow, reddish, blackish, or deep red-purple, reduced, protuberant, forming a platform, 1–2 mm, 5–25% as long as beak; teeth ascending, green, purple sometimes with reddish tip, yellow, or blackish, 0.5–1 mm. |
straight to slightly curved, 26–43 mm; tube 16–24 mm; beak exserted, adaxially green to yellow, 9–19 mm; abaxial lip whitish or dark green, reduced, not exserted, sometimes visible through front calyx cleft, 1.5–2 mm, 20% as long as beak, glabrous or sparsely hairy, hairs spreading; teeth reduced, incurved, deep green, 0.5–1 mm. |
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Calyces | proximally pale green, rarely pale purple, distally bright red or red-orange, rarely orange or yellow, or bright red or red-orange throughout, 20–42 mm; abaxial clefts 13–27 mm, adaxial 7–17 mm, abaxial 40–70% of calyx length, adaxial 12–33% of calyx length, deeper than laterals, lateral 0–7.5 mm, 0–35% of calyx length; lobes linear to narrowly triangular, strongly curved away from stem proximally and distally obviously curved toward stem, apex acute to acuminate to obtuse. |
colored as bracts, sometimes more deeply pigmented and contrasting with bract coloration, 24–31 mm; abaxial and adaxial clefts 12–19 mm, 45–65% of calyx length, deeper than laterals, lateral 7–11 mm, 25–32% of calyx length; lobes lanceolate, narrowly triangular, or almost oblong, apex acute, sometimes obtuse or acuminate. |
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2n | = 24, 48, 72, 96. |
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Castilleja subinclusa |
Castilleja chrymactis |
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Phenology | Flowering (Jan–)Feb–Sep(–Oct). | |||||||||
Habitat | Glacial outwash plains, river flats, moist or wet openings, beach meadows, roadsides. | |||||||||
Elevation | 0–20 m. (0–100 ft.) | |||||||||
Distribution |
CA; nw Mexico
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AK |
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Discussion | Varieties 3 (3 in the flora). Castilleja subinclusa is divided into three varieties with somewhat discontinuous ranges. Identification of the varieties is often difficult when comparing only a single trait but is more easily accomplished when the characters are considered in a suite. The three varieties are also separable by range and habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja chrymactis is restricted to the coast of southeastern Alaska, from Glacier Bay to Yakutat. In some respects, it resembles a hybrid swarm between C. miniata and C. unalaschcensis. If C. chrymactis originated as a hybrid, at present it has formed a number of populations and has a stable morphology that differs from both putative parents, especially in calyx structure. Reports of its introduction to Point Reyes, California (T. I. Chuang and L. R. Heckard 1993b), are based on C. leschkeana, a distinctive species known only from two records near the type locality and presumed to be extinct. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 657. | FNA vol. 17, p. 596. | ||||||||
Parent taxa | ||||||||||
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Name authority | Greene: Pittonia 4: 2. (1899) — (as Castilleia) | Pennell: Proc. Acad. Nat. Sci. Philadelphia 86: 539. (1934) | ||||||||
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