Castilleja scabrida |
Castilleja peckiana |
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rough Indian paintbrush, rough paintbrush |
peak paintbrush, Peck's Indian paintbrush, Peck's paintbrush |
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Habit | Herbs, perennial, 0.7–1.5(–2.2) dm; from a woody caudex; with a taproot or stout, branched roots. | Herbs, perennial, (1.8–)2.4–6 dm; from a woody caudex; with a stout taproot. | ||||
Stems | several, decumbent, often sprawling, distally ascending, unbranched unless injured, rarely with small, leafy axillary shoots, hairs spreading, whitish, short, ± stiff, eglandular. |
few to many, erect or ascending, often branched distally, sometimes unbranched, sometimes with short axillary shoots, proximal hairs retrorse to appressed, short to moderately long, distal hairs spreading, longer, soft, often mixed with short stipitate-glandular ones. |
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Leaves | gray-green, sometimes green or reddish purple, reduced and scalelike on proximal 10–25% of stem, linear, lanceolate, or narrowly elliptic, 1.5–4(–5) cm, not fleshy, margins plane, flat or involute, 0–3(–5)-lobed, apex acute; lobes spreading or spreading-ascending, linear or lanceolate, apex acute. |
green, linear-lanceolate, rarely broadly lanceolate, (1.2–)2.5–8(–9) cm, not fleshy, margins plane, flat or involute, 0(–3)-lobed, apex acute to rounded; lobes ascending-spreading, narrowly lanceolate to linear, apex acute or obtuse. |
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Inflorescences | 2.5–10 × 2.5–5 cm; bracts proximally green to deep greenish purple, distally bright red, sometimes brick red or orange-red, linear to lanceolate, 3–5(–7)-lobed; lobes spreading, linear-lanceolate to lanceolate, sometimes expanded near tip, long, arising near base and more distally, apex acute to obtuse. |
(2–)4–17 × 1.5–3 cm; bracts proximally greenish, distally red, orange-red, or orange, proximal sometimes lanceolate, distal broadly lanceolate to ovate, (0–)3(–7)-lobed; lobes spreading to ascending, linear to lanceolate, long, arising near or below mid length, central lobe apex rounded, lateral ones acute. |
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Corollas | straight or slightly curved, 25–40(–45) mm; tube 11–24 mm; beak exserted, adaxially green to yellow, 10–17(–20) mm; abaxial lip green, reduced, visible through front cleft, 1.5–2.5 mm, 10–15% as long as beak; teeth incurved, green, 0.5–1 mm. |
straight, 23–30(–35) mm; tube 12–20 mm; beak exserted, adaxially green, 8–12(–14) mm; abaxial lip pale to deep green, reduced, rounded, 0.5–1.5 mm, 10–20% as long as beak; teeth erect to incurved, green, (0.4–)0.7–1.2(–2) mm. |
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Calyces | colored as bracts, 18–33 mm; abaxial clefts 6–12 mm, adaxial 8–15 mm, clefts 40–60% of calyx length, deeper than laterals, lateral 2–6 mm, 15–25% of calyx length; lobes lanceolate or narrowly triangular, apex acute to acuminate. |
proximally pale yellow or greenish, distally colored as bracts, (15–)18–28 mm; abaxial and adaxial clefts 6–12 mm, 40–45% of calyx length, deeper than laterals, lateral 2.5–8 mm, 20–30% of calyx length; lobes linear to narrowly lanceolate, apex acute to acuminate. |
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2n | = 24. |
= 72, 96, ca. 120. |
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Castilleja scabrida |
Castilleja peckiana |
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Phenology | Flowering (Apr–)May–Aug. | |||||
Habitat | Open conifer forests, sagebrush slopes, riparian meadows, shores. | |||||
Elevation | 1400–2600 m. (4600–8500 ft.) | |||||
Distribution |
AZ; CO; NM; NV; UT
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ID; NV; OR
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Discussion | Varieties 2 (2 in the flora). Castilleja scabrida consists of two varieties distinguished by range and habitat differences, as well as by the morphological characters of the key. Both varieties are often confused with the similar and broadly sympatric C. chromosa but can be differentiated from the latter most easily by the presence of scalelike vestigial leaves on the proximal stems of C. scabrida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
When describing Castilleja peckiana, Pennell noted that its variation approached C. hispida on one hand and C. miniata on the other, and it is likely of allopolyploid derivation. L. R. Heckard (1968) found chromosome numbers of n = 36, 48, and ca. 60. He hypothesized that C. chromosa, C. hispida var. acuta, and C. miniata were likely involved in its ancestry, and possibly C. pruinosa as well. Heckard suggested subsequent introgression among the derived forms introduced further complexity. Though complex, these forms are self-perpetuating and appear morphologically stable within their range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 654. | FNA vol. 17, p. 643. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Eastwood: Bull. Torrey Bot. Club 29: 523. (1902) — (as Castilleia) | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 9. (1941) | ||||
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