Castilleja rupicola |
Castilleja lacera |
|
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cliff Indian paintbrush, cliff paintbrush |
cut-leaf owl's clover, cut-leaf paintbrush, cutleaf Indian paintbrush, foothill owl's clover |
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Habit | Herbs, perennial, (0.8–)1–2(–3) dm; from a woody caudex; with a taproot. | Herbs, annual, 0.5–4 dm; with fibrous roots. |
Stems | many, decumbent to ascending, unbranched, sparsely pubescent, hairs spreading, wavy, fairly short, soft, eglandular, sometimes glabrous proximally. |
solitary, erect, unbranched or branched, hairs spreading, long, soft, scattered among more numerous, medium length, stipitate-glandular ones. |
Leaves | purple to green, narrowly, rarely broadly, lanceolate, 1.4–4 cm, not fleshy, margins plane, flat to involute, (0–)3–5(–7)-lobed, apex acute to acuminate; lobes divergent, spreading-ascending, linear, long, not much narrower than mid blade, often with secondary lobes, creating little frilly fans, apex acute or obtuse. |
green or purplish, linear to narrowly lanceolate, 1–5 cm, not fleshy, margins plane, flat, 0–5(–7)-lobed, apex acuminate; lobes spreading to ascending, linear, apex acuminate to acute. |
Inflorescences | 2–6 × 2–3.5 cm; bracts proximally greenish or deep purple near base, distally red, scarlet, or crimson to red-orange, rarely orange, salmon, pink, or yellowish white, ovate to orbicular in outline, 5(–9)-lobed; lobes spreading, linear to linear-lanceolate, long, arising below mid length, apex acute to rounded. |
(1.5–)3–14 × 2–3 cm; bracts green throughout, sometimes proximally green, distally white on apices, lanceolate to ovate, 3–7-lobed; lobes spreading to ascending, linear to narrowly lanceolate, long, arising below mid length, apex obtuse to acute. |
Corollas | straight or slightly curved, 25–35(–45) mm; tube 9–15 mm; beak exserted, adaxially green, purplish, or yellow-green, 14–22 mm; abaxial lip deep green, reduced, 0.5–2 mm, 6–12% as long as beak; teeth incurved to erect, green, 0.5 mm. |
straight, 10–22 mm; tube 8–15 mm; abaxial lip and beak exserted; beak adaxially yellow to greenish, 3–6 mm, densely puberulent; abaxial lip yellow with purple dots at base, inflated, pouches 3, central pouch slightly 2-lobed, pouches 4–8 mm wide, 3–6 mm deep, side pouches curving up a little at tip, 2–5 mm, 75–95% as long as beak; teeth erect, white or yellow, 0.5–2 mm. |
Calyces | proximally purple, green, or whitish, distally colored as bract lobes, 15–25 mm; abaxial and adaxial clefts 8 mm, ca. 40–50% of calyx length, deeper than laterals, lateral 1–5 mm, 10–20% of calyx length; lobes triangular, apex obtuse or acute. |
light green, lobes green, 7–13 mm; abaxial and adaxial clefts 3.5–8 mm, 50–67% of calyx length, lateral 2.5–5 mm, ca. 40% of calyx length; lobes narrowly to broadly lanceolate, apex acute to acuminate. |
Stigmas | equal to or slightly exserted from beak. |
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2n | = 22, 24. |
|
Castilleja rupicola |
Castilleja lacera |
|
Phenology | Flowering (May–)Jun–Sep. | Flowering (Mar–)Apr–Jul(–Aug). |
Habitat | Sunny rocky slopes, scree, talus, ledges, fellfields, subalpine to alpine. | Grasslands, meadows, moist flats, vernal pool margins, moist forest openings, serpentine slopes and ledges, roadsides. |
Elevation | (200–)1000–2500 m. ((700–)3300–8200 ft.) | 0–2700 m. (0–8900 ft.) |
Distribution |
OR; WA; BC
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CA; OR
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Discussion | Castilleja rupicola is usually found in the subalpine and lower alpine zones in the Cascade Range from extreme southern British Columbia south to northern Douglas County, Oregon. Though it can be numerous where it occurs, the species as a whole is uncommon. One atypical population occurs in a moist, shaded, mossy, north-facing ravine on the Oregon side of the Columbia River Gorge, at less than 250 m. These plants often bear secondary divisions on deeply dissected leaves and bracts. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja lacera is found in a wide range of elevations in the central and northern Sierra Nevada region and in the Siskiyou Mountains region of northwestern California and southwestern Oregon. Reports from the Coast Ranges north of the San Francisco Bay region and south of the Siskiyou region in western California are referable to other yellow-flowered annuals, including C. ambigua, C. rubicundula var. lithospermoides, Triphysaria eriantha subsp. eriantha, and T. versicolor subsp. faucibarbata. Although most similar to C. rubicundula, C. lacera is somewhat smaller in stature and flower size. It is also easily confused with yellow-flowered populations of C. tenuis, which has smaller flowers and an included stigma. Two chromosome numbers are known for this species, the more northern populations being diploid, and those to the south having an apparently aneuploid count of 2n = 22, which is unique in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 653. | FNA vol. 17, p. 617. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | Orthocarpus lacerus | |
Name authority | Piper: Erythea 6: 45. (1898) — (as Castilleia) | (Bentham) T. I. Chuang & Heckard: Syst. Bot. 16: 657. (1991) |
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