Castilleja rubida |
Castilleja gleasoni |
|
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little reddish Indian paintbrush, purple alpine paintbrush, purple paintbrush, Wallowa alpine paintbrush |
Mount Gleason paintbrush, Mt. Gleason paintbrush |
|
Habit | Herbs, perennial, 0.5–1.5 dm; from a woody caudex; with a taproot. | Herbs or subshrubs, perennial, 3–8 dm; from a woody caudex; with a taproot. |
Stems | several, decumbent, or ascending, unbranched, hairs moderately dense, spreading, short and long, soft, eglandular and glandular. |
several to many, erect to ascending, branched, sometimes unbranched, hairs moderately dense, spreading, ± matted, ash gray, branched, sometimes unbranched, medium length, soft, often mixed with shorter, unbranched, stipitate-glandular ones, not obscuring surface. |
Leaves | green to purple, linear to narrowly lanceolate, 0.7–3.2 cm, not fleshy, margins plane, slightly involute, 3–5-lobed, apex narrowly acute to acuminate; lobes ascending-spreading, narrowly linear to filiform, often curling, often short, apex acute or obtuse. |
ash gray, linear-lanceolate to narrowly oblong or narrowly oblanceolate, 2–6 cm, not fleshy, margins wavy, involute, 0(–3)-lobed, apex acute to obtuse; lobes spreading, narrowly lanceolate, apex acute to obtuse. |
Inflorescences | 2.5–6 × 1–2 cm; bracts purple, deep burgundy, or lavender throughout, rarely pink or yellowish white throughout, sometimes pink or dull whitish on distal margins and apices, oblong, 3–5(–7)-lobed; lobes spreading, linear, medium length, proximal lobes arising below mid length, center lobe apex rounded to obtuse, lateral ones acute to obtuse. |
10–15(–30) cm; bracts proximally greenish to dull brownish purple, distally red to deep red or red-orange, lanceolate to oblong or narrowly ovate, (0–)3(–5)-lobed; lobes ascending to spreading, narrowly oblong to narrowly lanceolate, medium length, arising below mid length, center lobe apex obtuse or toothed, lateral ones acute. |
Corollas | straight, 12–15 mm; tube 14–16 mm; abaxial lip and beak exserted; beak adaxially green, 5–6 mm; abaxial lip colored as distal portion of bracts, prominent, pouches 3, central one grooved, pouches not strongly inflated, 4–5 mm, 80–100% as long as beak; teeth erect, appressed to beak, colored as distal portions of bracts, 1.5–2.5 mm. |
± straight, 18–30 mm; tube 9–15 mm; beak exserted, adaxially yellow, 9–15(–20) mm; abaxial lip spreading, deep green to ± black, reduced, 0.5–2 mm, 20% as long as beak; teeth incurved, green, 0.2–1 mm. |
Calyces | colored as bracts, 10–12 mm; all 4 clefts subequal, 3.5–6.5 mm, 35–55% of calyx length; lobes broadly linear or linear-triangular, apex obtuse to acute. |
proximally pale to green, sometimes dull purple, often paler or greener than bracts, distal 1/4 or less colored as bract lobes, 12–17 mm; abaxial and adaxial clefts 3–8 mm, 40% of calyx length, deeper than laterals, lateral 0.5–3 mm, 20–25% of calyx length; lobes ovate to narrowly triangular, apex obtuse to acute. |
Stigmas | green. |
|
2n | = 72. |
|
Castilleja rubida |
Castilleja gleasoni |
|
Phenology | Flowering Jul–Aug. | Flowering May–Jun. |
Habitat | Rocky slopes, ledges, dry to moist gravelly flats and ridges, alpine, limestone, rarely on river cobbles at lower elevations. | Ledges, rocky slopes, open yellow pine forests, montane chaparral or sagebrush. |
Elevation | 2200–3000 m. (7200–9800 ft.) | 900–2200 m. (3000–7200 ft.) |
Distribution |
OR |
CA |
Discussion | Castilleja rubida is a rare alpine species endemic to a few limestone peaks in the Wallowa Mountains of northeastern Oregon, entirely within the Eagle Cap Wilderness Area. It is likely derived from the C. nana complex, found in the mountains of eastern California and Nevada, but it is amply distinct. Due to its very limited range and small population numbers, C. rubida is a species of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja gleasoni is an uncommon plant endemic to the upper elevations of the San Gabriel Mountains of Los Angeles County. It was treated as a polyploid derivative of C. affinis var. affinis (as subsp. affinis) × C. foliolosa (T. I. Chuang and L. R. Heckard 1993b), a hypothetical ancestry supported by a chromosome number of 2n = 72. Others have placed it as a subspecies of C. pruinosa, a similar species. However, the morphology of C. gleasoni also suggests it could have originated as a cross between the diploid C. foliolosa and a tetraploid form of C. martini; careful morphological and molecular analyses are needed to determine its true ancestry. Castilleja gleasoni is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 653. | FNA vol. 17, p. 610. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. pruinosa subsp. gleasoni | |
Name authority | Piper: Bull. Torrey Bot. Club 27: 398. (1900) — (as Castilleia) | Elmer: Bot. Gaz. 39: 51. (1905) — (as Castilleia) |
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