Castilleja rhexiifolia |
Castilleja pruinosa |
|
---|---|---|
alpine paintbrush, rhexia-leaf paintbrush, Rocky Mountain paintbrush, rosy paintbrush, split-leaf Indian paintbrush |
frosted Indian paintbrush, frosted paintbrush, pruinose paintbrush |
|
Habit | Herbs, perennial, (1–)2.5–6(–8) dm; from a woody caudex; with a taproot. | Herbs to subshrubs, perennial, (1.2–)2–8 dm; from a woody caudex; with a taproot. |
Stems | few to several, erect or ascending, unbranched, sometimes branched, proximally glabrous to glabrate, distally sparsely hairy, hairs spreading, ± long, soft, eglandular, sometimes stipitate-glandular. |
several to many, erect or ascending, branched or unbranched, sometimes with inconspicuous, short, leafy axillary shoots, hairs ± dense, spreading to appressed, whitish or ashy gray, long, soft, much-branched, rarely unbranched, eglandular or obscurely glandular. |
Leaves | green or purple-tinged, linear, narrowly to broadly lanceolate, oblong, or ovate, 3–6(–7) cm, not fleshy, margins plane, flat to slightly involute, prominently veined, 0(–3)-lobed, apex acute to rarely obtuse; lobes ascending or spreading, linear to filiform, much smaller than mid blade, short, apex acute. |
green, linear-lanceolate to narrowly or broadly lanceolate, 2–8 cm, not fleshy, margins plane to sometimes wavy, flat or involute, 0-lobed, sometimes 3–5-lobed distally, apex obtuse to rounded, sometimes acute; lobes sometimes divergent, ascending-spreading, linear, apex acute. |
Inflorescences | 2.5–15 × 1.5–4.5 cm; bracts pink-purple, red-purple, purple, or crimson throughout, rarely reddish, yellowish, or white throughout, or proximally greenish, dull brownish purple, or deep purple, distally colored as above, broadly lanceolate to ovate or obovate, (0–)3–5(–7)-lobed; lobes ascending to erect, linear or lanceolate-acuminate, short, arising near or above mid length, central lobe apex obtuse to broadly rounded, lateral ones acute to sometimes rounded. |
3–20 × 1.5–4 cm; bracts proximally greenish, distally abruptly red, sometimes yellow, salmon, or orange-red, often with a narrow yellow medial band, lanceolate, broadly lanceolate, or oblong, 0–5-lobed; lobes spreading to ascending, linear, long, arising near mid length, apex obtuse to acute. |
Corollas | straight, 15–30(–36) mm; tube (11–)12–22(–24) mm; beak exserted, adaxially green, yellow, or tinged with red, 7–12 mm; abaxial lip deep green, reduced, 1.5–3.5 mm, to 33% as long as beak; teeth incurved, green or white, 0.5–2 mm. |
slightly curved, 20–51 mm; tube 12–21 mm; beak long-exserted, adaxially green, yellow, or red, 12–33 mm; abaxial lip deep green, reduced, 1–2.5 mm, ca. 10% as long as beak; teeth incurved to erect, white, yellow, or green, 0.5–2 mm. |
Calyces | proximally green, purplish, or whitish, distally colored as bracts, 15–25 mm; abaxial and adaxial clefts 8–12(–15) mm, 40–50% of calyx length, deeper than laterals, lateral 2–5(–8) mm, 15–25(–30)% of calyx length; lobes oblong to triangular or ovate, apex obtuse to rounded, sometimes acute. |
proximally green to yellow, distally and abruptly colored as in distal bracts, often with a central yellow band, 13–28 mm; abaxial clefts 5–13 mm, adaxial 8–14 mm, clefts 33–70% of calyx length, deeper than laterals, lateral 3–9 mm, 20–25% of calyx length; lobes lanceolate to triangular, apex acute to acuminate. |
2n | = 24, 48, 96. |
= 48. |
Castilleja rhexiifolia |
Castilleja pruinosa |
|
Phenology | Flowering (Mar–)May–Sep. | Flowering Mar–Aug. |
Habitat | Moist meadows, open forests, slopes and ridges, sun or shade, subalpine to alpine. | Dry flats, rocky slopes, talus, thickets, open forests and forest edges, usually over serpentine. |
Elevation | 1800–4000 m. (5900–13100 ft.) | 50–2600 m. (200–8500 ft.) |
Distribution |
CO; ID; MT; NM; OR; UT; WA; WY; AB; BC
|
CA; OR
|
Discussion | Castilleja rhexiifolia is a characteristic and common paintbrush in the Rocky Mountains region, from central Alberta to northern New Mexico. Disjunct populations also occur in northeastern Oregon, in the Blue, Ochoco, and Wallowa mountains, as well as near the summit of Abercrombie Mountain in the Selkirk Mountains of northeastern Washington. The inflorescence colors are variable. Typical plants are red-purple. However, many other shades of pink-white, pink-purple, and crimson are common as well. Many populations contain a wide range of color variants. Intergradation is occasionally seen with C. miniata, C. occidentalis, and C. septentrionalis. However, for the most part, these four species remain distinct, and their ranges are not entirely overlapping. In the Wallowa Mountains, a recurrent and variable hybrid form between C. chrysantha and C. rhexiifolia was described as C. wallowensis Pennell. In the north Cascade Range of Washington, plants of C. elmeri with crimson inflorescences are sometimes identified as C. rhexiifolia. In Glacier National Park, Montana, in the vicinity of Logan Pass, C. rhexiifolia frequently hybridizes with C. occidentalis, creating a number of hybrid morphologies. Both of the parents are common in the area. Similar hybrids are also found in the Canadian Rocky Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja pruinosa is a common and variable paintbrush at moderate elevations in the southern half of the Cascade Range, occurring westward in Oregon and in adjacent northwestern California. It is also found in the northern Sierra Nevada. The often dense pubescence of branched hairs distinguishes it from other similar species within its range. Castilleja gleasoni was sometimes included within C. pruinosa, but C. gleasoni is likely of hybrid origin and occurs only in the San Gabriel Mountains of southern California. There are significant morphological differences between C. gleasoni and C. pruinosa. Castilleja pruinosa forms occasional hybrids with C. applegatei var. pinetorum and with C. brevilobata. In some populations in Douglas and eastern Lane counties, Oregon, C. pruinosa forms introgressive swarms with C. hispida var. hispida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 651. | FNA vol. 17, p. 647. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. oregonensis | |
Name authority | Rydberg: Mem. New York Bot. Gard. 1: 356. (1900) — (as rhexifolia) | Fernald: Erythea 6: 50. (1898) — (as Castilleia) |
Web links |