Castilleja puberula |
Castilleja peckiana |
|
---|---|---|
alpine paintbrush, short-flower paintbrush, shortflower Indian paintbrush |
peak paintbrush, Peck's Indian paintbrush, Peck's paintbrush |
|
Habit | Herbs, perennial, 0.8–1.5 dm; from a woody caudex; with a taproot. | Herbs, perennial, (1.8–)2.4–6 dm; from a woody caudex; with a stout taproot. |
Stems | few to several, erect or ascending, unbranched, sometimes branched, hairs retrorse, short, soft, eglandular. |
few to many, erect or ascending, often branched distally, sometimes unbranched, sometimes with short axillary shoots, proximal hairs retrorse to appressed, short to moderately long, distal hairs spreading, longer, soft, often mixed with short stipitate-glandular ones. |
Leaves | green to purple, linear to linear-lanceolate, 2–3.3 cm, not fleshy, margins plane, involute, 0–3(–5)-lobed, apex acuminate; lobes ascending-spreading to widely spreading, narrowly linear, apex acute. |
green, linear-lanceolate, rarely broadly lanceolate, (1.2–)2.5–8(–9) cm, not fleshy, margins plane, flat or involute, 0(–3)-lobed, apex acute to rounded; lobes ascending-spreading, narrowly lanceolate to linear, apex acute or obtuse. |
Inflorescences | 4–5 × 1.5–2.5 cm; bracts greenish throughout, or proximally greenish, distally bright yellow, yellow-green, or yellow-orange on apices, narrowly lanceolate to linear-lanceolate, 0–5(–7)-lobed; lobes spreading, filiform to linear, long, proximal lobes arising in proximal 1/2, apex acute to acuminate. |
(2–)4–17 × 1.5–3 cm; bracts proximally greenish, distally red, orange-red, or orange, proximal sometimes lanceolate, distal broadly lanceolate to ovate, (0–)3(–7)-lobed; lobes spreading to ascending, linear to lanceolate, long, arising near or below mid length, central lobe apex rounded, lateral ones acute. |
Corollas | straight or slightly curved, 18–21 mm; tube 13–15 mm; beak exserted, adaxially yellow to yellow-green, 6–8 mm; abaxial lip green, reduced, visible in front cleft, slightly pouched, 2.5–3.5 mm, 35–60% as long as beak; teeth erect, yellow, 1 mm. |
straight, 23–30(–35) mm; tube 12–20 mm; beak exserted, adaxially green, 8–12(–14) mm; abaxial lip pale to deep green, reduced, rounded, 0.5–1.5 mm, 10–20% as long as beak; teeth erect to incurved, green, (0.4–)0.7–1.2(–2) mm. |
Calyces | light green to yellowish, margins sometimes yellow, 10–17 mm; abaxial clefts 8–9 mm, adaxial 2–3 mm, abaxial ca. 50% of calyx length, adaxial ca. 20% of calyx length, deeper than laterals, lateral 1.9–2.5 mm, 12–20% of calyx length; lobes narrowly triangular, apex acute to acuminate. |
proximally pale yellow or greenish, distally colored as bracts, (15–)18–28 mm; abaxial and adaxial clefts 6–12 mm, 40–45% of calyx length, deeper than laterals, lateral 2.5–8 mm, 20–30% of calyx length; lobes linear to narrowly lanceolate, apex acute to acuminate. |
2n | = 24. |
= 72, 96, ca. 120. |
Castilleja puberula |
Castilleja peckiana |
|
Phenology | Flowering (Feb–)Jul–Aug. | Flowering (Apr–)May–Aug. |
Habitat | Moist meadows, stream banks, mesic rocky slopes, tundra, subalpine and alpine. | Open conifer forests, sagebrush slopes, riparian meadows, shores. |
Elevation | 2700–3900 m. (8900–12800 ft.) | 1400–2600 m. (4600–8500 ft.) |
Distribution |
CO; MT
|
ID; NV; OR
|
Discussion | Castilleja puberula is a near-endemic of alpine communities in the Rocky Mountains of Colorado in Boulder, Clear Creek, Gilpin, Grand, Larimer, and Park counties. A highly disjunct population occurs near the summit of Mt. Jefferson on the Continental Divide in Beaverhead County, Montana. Castilleja puberula appears to be a high-elevation isolate from the widespread C. flava, which is common at lower elevations in the Intermountain Region. A single case of hybridization with C. septentrionalis is known from Clear Creek County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
When describing Castilleja peckiana, Pennell noted that its variation approached C. hispida on one hand and C. miniata on the other, and it is likely of allopolyploid derivation. L. R. Heckard (1968) found chromosome numbers of n = 36, 48, and ca. 60. He hypothesized that C. chromosa, C. hispida var. acuta, and C. miniata were likely involved in its ancestry, and possibly C. pruinosa as well. Heckard suggested subsequent introgression among the derived forms introduced further complexity. Though complex, these forms are self-perpetuating and appear morphologically stable within their range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 648. | FNA vol. 17, p. 643. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Name authority | Rydberg: Bull. Torrey Bot. Club 31: 644. (1905) | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 9. (1941) |
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