Castilleja pruinosa |
Castilleja martini |
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frosted Indian paintbrush, frosted paintbrush, pruinose paintbrush |
Camp Martin paintbrush |
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Habit | Herbs to subshrubs, perennial, (1.2–)2–8 dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.5–5(–8) dm; from a woody caudex; with a taproot. | ||||
Stems | several to many, erect or ascending, branched or unbranched, sometimes with inconspicuous, short, leafy axillary shoots, hairs ± dense, spreading to appressed, whitish or ashy gray, long, soft, much-branched, rarely unbranched, eglandular or obscurely glandular. |
several to many, erect or ascending, sometimes curved at base, unbranched or branched, sometimes with short, leafy axillary shoots, hairs spreading, short or long and distally short, stiff or soft, stipitate-glandular. |
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Leaves | green, linear-lanceolate to narrowly or broadly lanceolate, 2–8 cm, not fleshy, margins plane to sometimes wavy, flat or involute, 0-lobed, sometimes 3–5-lobed distally, apex obtuse to rounded, sometimes acute; lobes sometimes divergent, ascending-spreading, linear, apex acute. |
green to sometimes purple, narrowly oblong or lanceolate to sometimes linear or narrowly ovate, (0.8–)1–5(–6.5) cm, not fleshy, margins wavy, flat or involute, 0–5-lobed, apex rounded or obtuse, rarely acute; lobes ascending-spreading or divaricate, linear to narrowly oblong, apex acute. |
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Inflorescences | 3–20 × 1.5–4 cm; bracts proximally greenish, distally abruptly red, sometimes yellow, salmon, or orange-red, often with a narrow yellow medial band, lanceolate, broadly lanceolate, or oblong, 0–5-lobed; lobes spreading to ascending, linear, long, arising near mid length, apex obtuse to acute. |
2.5–25 × 1.5–4 cm; bracts proximally greenish, distally red, orange-red, or pale orange, rarely yellow, lanceolate or broadly lanceolate to oblong, (0–)3–5(–7)-lobed, plus 1–3 shallow teeth on apex of central lobe (var. clokeyi); lobes ascending or spreading, linear, lanceolate, or oblanceolate, medium length or long, arising at or near mid length, central lobe apex obtuse to rounded, lateral ones acute. |
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Corollas | slightly curved, 20–51 mm; tube 12–21 mm; beak long-exserted, adaxially green, yellow, or red, 12–33 mm; abaxial lip deep green, reduced, 1–2.5 mm, ca. 10% as long as beak; teeth incurved to erect, white, yellow, or green, 0.5–2 mm. |
straight to slightly curved, (19–)22–45 mm; tube 11–22 mm; abaxial lip rarely exserted, beak exserted; beak adaxially green to yellowish, 9–21(–23) mm; abaxial lip green or deep green with white teeth or red with green teeth, reduced, not inflated, often visible through front cleft, 1–2 mm, ca. 10% as long as beak; teeth incurved, green to white, 0.5–1 mm. |
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Calyces | proximally green to yellow, distally and abruptly colored as in distal bracts, often with a central yellow band, 13–28 mm; abaxial clefts 5–13 mm, adaxial 8–14 mm, clefts 33–70% of calyx length, deeper than laterals, lateral 3–9 mm, 20–25% of calyx length; lobes lanceolate to triangular, apex acute to acuminate. |
proximally pale green, green, or whitish, sometimes with a yellow central band, distally colored as bracts, 14–26(–28) mm; abaxial and adaxial clefts 3–10 mm, 20–50% of calyx length, deeper than laterals, lateral 1–5 mm, 15–20% of calyx length; lobes lanceolate to broadly lanceolate, oblong, or ovate, apex acute, obtuse, or rounded. |
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2n | = 48. |
= 24, 48, 72. |
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Castilleja pruinosa |
Castilleja martini |
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Phenology | Flowering Mar–Aug. | |||||
Habitat | Dry flats, rocky slopes, talus, thickets, open forests and forest edges, usually over serpentine. | |||||
Elevation | 50–2600 m. (200–8500 ft.) | |||||
Distribution |
CA; OR
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CA; NV; nw Mexico |
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Discussion | Castilleja pruinosa is a common and variable paintbrush at moderate elevations in the southern half of the Cascade Range, occurring westward in Oregon and in adjacent northwestern California. It is also found in the northern Sierra Nevada. The often dense pubescence of branched hairs distinguishes it from other similar species within its range. Castilleja gleasoni was sometimes included within C. pruinosa, but C. gleasoni is likely of hybrid origin and occurs only in the San Gabriel Mountains of southern California. There are significant morphological differences between C. gleasoni and C. pruinosa. Castilleja pruinosa forms occasional hybrids with C. applegatei var. pinetorum and with C. brevilobata. In some populations in Douglas and eastern Lane counties, Oregon, C. pruinosa forms introgressive swarms with C. hispida var. hispida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja martini is sometimes treated as a subspecies of C. applegatei, but there are relatively consistent morphological discontinuities between these two variable complexes. See notes under 4. C. applegatei. Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 647. | FNA vol. 17, p. 625. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
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Subordinate taxa | ||||||
Synonyms | C. applegatei subsp. martini | |||||
Name authority | Fernald: Erythea 6: 50. (1898) — (as Castilleia) | Abrams: Bull. S. Calif. Acad. Sci. 1: 69. (1902) | ||||
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