Castilleja plagiotoma |
Castilleja parviflora |
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Mojave desert paintbrush, Mojave Indian paintbrush, Mojave paintbrush |
magenta paintbrush, mountain Indian paintbrush, mountain or rosy or small-flower paintbrush, small-flower paintbrush |
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Habit | Herbs, perennial, 2.3–5.3(–6) dm; from a woody caudex; with a taproot. | Herbs, perennial, (0.6–)1–4(–5) dm; from a woody caudex; with a taproot or stout, branched roots. | ||||||||||||
Stems | several, erect to weakly ascending, often leaning on nearby shrubs, much-branched, often with short, leafy axillary shoots, hairs sparse, spreading, short or long, soft, branched or unbranched, eglandular. |
several or many, erect or ascending, unbranched except for short, leafy axillary shoots, glabrate proximally, hairy distally, hairs sparse, spreading, ± matted, long, soft, minute-glandular. |
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Leaves | gray-green or green to purple to ± deep red, linear or linear-lanceolate, 2–5 cm, not fleshy, margins plane to ± wavy, flat or involute, 3–5(–7)-lobed, apex acuminate to acute; lobes spreading-ascending, sometimes widely so, narrowly linear, apex acute or obtuse. |
green or gray-green to purple-tinged or deep purple, often blackening on drying, narrowly to broadly lanceolate or elliptic, rarely linear, 1.5–5 cm, not fleshy, margins plane, sometimes ± wavy, flat, (0–)3–9-lobed, apex acute to acuminate or obtuse; lobes spreading or ascending, linear, sometimes lanceolate, much narrower than terminal lobe, evenly spaced, short, apex acute. |
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Inflorescences | 3–20 × 0.5–1.5 cm; bracts green throughout, sometimes tinged dull purple, proximal sometimes linear-lanceolate, distal or all bracts oblong, 3–5(–7)-lobed; lobes spreading to ascending, linear to oblanceolate, short to long, proximal lobes arising near mid length, sometimes wavy-margined, central lobe apex rounded to truncate, lateral ones obtuse to rounded. |
2–16 × 1–3.5 cm; bracts proximally greenish, dull, deep purple, or reddish purple, distally pink, pink-purple, magenta, deep rose, crimson, cream, or white, sometimes red, pale orange, or red-orange, lanceolate to broadly elliptic or ovate, 3–7-lobed; lobes spreading to ascending, linear, lanceolate, or lanceolate-acuminate, short to medium length, arising at or near mid length, apex obtuse to acute, central lobes sometimes rounded. |
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Corollas | straight, 13–20 mm; tube 5–7 mm; beak short-exserted, sometimes shorter than abaxial calyx segments, adaxially yellow, rarely greenish, 7–10 mm; abaxial lip pale green to yellowish, reduced, 0.5–1.5 mm, 7–17% as long as beak; teeth reduced to apiculations, pale green to yellowish, 0.5–0.8 mm. |
straight or slightly curved, 12–30 mm; tube 8–19 mm; beak exserted or subequal to calyx, adaxially green-yellowish or red, 5.5–11 mm; abaxial lip green, brown, or yellow, sometimes purple, reduced, slightly or not inflated and pouched, 1–3 mm, 20–45% as long as beak; teeth erect, green, white, yellow, pink, or red, 0.5–2 mm. |
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Calyces | light green, cream, or light yellow, often appearing white from dense white-woolly pubescence, 10–18 mm; abaxial clefts 5–6.5 mm, closed by intertwined hairs and appearing 2 mm deep, adaxial 2.2–4.2 mm, abaxial ca. 25% of calyx length, adaxial ca. 20% of calyx length, shallower (or appearing much shallower), than laterals, lateral 4.6–7.5 mm, ca. 50% of calyx length; lobes: abaxial segments broad, paddle-shaped, 7 mm, inner surface of abaxial segments densely white-woolly, apex rounded to broadly obtuse, adaxial narrowly triangular, 5 mm, apex acute. |
colored as bracts, 12–28 mm; abaxial and adaxial clefts 6–15 mm, 40–70% of calyx length, deeper than laterals, lateral 1–8 mm, 10–35% of calyx length; lobes narrowly to broadly triangular, sometimes distally expanded and flaring, petaloid, apex obtuse or acute, sometimes rounded. |
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2n | = 24, 48. |
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Castilleja plagiotoma |
Castilleja parviflora |
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Phenology | Flowering Mar–Jul. | |||||||||||||
Habitat | Dry flats, rocky, sandy, or clayey slopes, ridges, sagebrush steppes, chaparral, desert scrub, pinyon woodlands. | |||||||||||||
Elevation | 200–2500 m. (700–8200 ft.) | |||||||||||||
Distribution |
CA
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AK; OR; WA; AB; BC; YT
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Discussion | Castilleja plagiotoma is unique in the genus in the structure of its calyces, with the lateral clefts deeper than the median clefts. The relative lengths of the calyx lobes are also unique, with the abaxial lobes exceeding the adaxial lobes by 2–3 mm. The abaxial lobes also bear a dense indument of whitish, branched hairs. The uniformly greenish bracts are uncommon among perennial species of Castilleja. Castilleja plagiotoma is scattered in the western Mojave Desert and the hills adjacent to the southwestern San Joaquin Valley, California. While not of immediate conservation concern, this species is uncommon and increasingly threatened by recreational vehicles, livestock grazing, residential development, and resource extraction. Castilleja plagiotoma is most often associated with and is likely parasitic on Eriogonum fasciculatum var. polifolium, but it is also often observed with Artemisia tridentata or other species. It is a known larval host plant for the butterfly, Euphydryas editha subsp. erlichii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Castilleja parviflora is a complex, geographically widespread, and often misunderstood species ranging from southeastern Alaska through much of British Columbia, southwestern Yukon, and the Rocky Mountains of extreme western Alberta and southward in the Cascade Range to central Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 646. | FNA vol. 17, p. 640. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Name authority | A. Gray: Proc. Amer. Acad. Arts 19: 93. (1883) — (as Castilleia) | Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, sér. 6, Sci. Math. 2(2): 158. (1832) | ||||||||||||
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