Castilleja oresbia |
Castilleja cusickii |
|
|---|---|---|
|
pale paintbrush, pale Wallowa Indian paintbrush, pale Wallowa paintbrush |
Cusick's Indian paintbrush, Cusick's paintbrush |
|
| Habit | Herbs, perennial, 0.9–3 dm; from a woody caudex; with a stout taproot. | Herbs, perennial, (1–)1.5–5(–6) dm; from a small, woody caudex; with a taproot or stout, branched roots. |
| Stems | few to several, erect or ascending, sometimes decumbent at base, unbranched or branched, hairs usually retrorse, medium length, ± soft, eglandular, mixed with very short-glandular ones, sometimes with spreading, long, soft ones. |
solitary or few to several, erect, often decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, eglandular, mixed with shorter, sometimes stiff, stipitate-glandular ones. |
| Leaves | green to purple, linear to lanceolate, 2–7 cm, not fleshy, margins plane, involute, 3–5(–7)-lobed, apex acuminate to acute; lobes spreading, linear to sometimes narrowly lanceolate, apex acute. |
green, sometimes with prominent red-purple veins, lanceolate-linear to broadly lanceolate, 2.5–4.5(–7) cm, not fleshy, margins plane, flat, (0–)3–5(–9)-lobed, apex acuminate to acute; lobes spreading-ascending, narrowly lanceolate, arising at or above mid length, apex acute. |
| Inflorescences | 2.5–18 × 1–3.5 cm; bracts pale green to yellow-green or pale, dull reddish brown throughout, or proximally so colored but changing gradually to cream or yellowish on distal margins, narrowly to broadly lanceolate, (3–)5–7(–9)-lobed; lobes ascending, linear, long, proximal lobes arising below mid length, central lobe apex obtuse, others acute. |
3.5–26 × 1.5–3.5 cm; bracts pale green, pale greenish yellow, or pale yellow throughout, or proximally pale green, pale greenish yellow, pale yellow, or reddish purple, distally white, yellow, pink, dull purple, or dull reddish purple, sometimes with a purple band below that, sometimes with veins darker than background color, lanceolate to oblong, 0–5(–9)-lobed; lobes ascending or spreading, linear to narrowly lanceolate, often short, arising above mid length, central lobe apex rounded, lateral ones acute to rounded. |
| Corollas | straight, 21–36 mm; tube 16–20 mm; teeth of abaxial lip often exserted, beak exserted; beak adaxially green, 4.2–5.5 mm; abaxial lip green to purple, distally white, conspicuous, slightly but noticeably pouched, often visible through front cleft, 3–5 mm, 67–100% as long as beak, puberulent; teeth erect, white, 1.8–2.1 mm. |
straight, 19–25(–28) mm; tube (13–)15–20 mm; abaxial lip included, beak not or slightly exserted; beak adaxially green to yellow-green, 4–7 mm; abaxial lip yellow, prominent, inflated, 2–4.5 mm, ca. 75% as long as beak; teeth incurved to erect, white, 1.3–2.1 mm. |
| Calyces | colored as bracts, 10–25 mm; abaxial and adaxial clefts 6–7 mm, 30–60% of calyx length, deeper than laterals, lateral 5–10 mm, 40–50% of calyx length; lobes linear, apex acute. |
green, pale green, or pale yellow, lobes yellow to pale yellow, 20–30 mm; abaxial and adaxial clefts 6.6–14 mm, 33–50% of calyx length, deeper than laterals, lateral 0–4(–4.5) mm, 0–15% of calyx length; lobes broadly rounded to triangular, apex obtuse to rounded. |
| 2n | = 24. |
|
Castilleja oresbia |
Castilleja cusickii |
|
| Phenology | Flowering May–Aug. | Flowering Apr–Aug. |
| Habitat | Dry slopes and plains, sagebrush meadows, grasslands, openings in conifer forests. | Moist meadows, swales, grasslands in sagebrush steppes, occasionally to subalpine. |
| Elevation | 900–2200 m. [3000–7200 ft.] | 500–2500 m. [1600–8200 ft.] |
| Distribution |
ID; OR
|
ID; MT; NV; OR; WA; WY; AB; BC
|
| Discussion | Castilleja oresbia is endemic to eastern Oregon and adjacent Idaho. It is easily confused with both varieties of C. pallescens, which also occur in sagebrush habitats. Castilleja oresbia has longer calyx lobes and softer pubescence than C. pallescens var. pallescens, although some transitional specimens are found. Castilleja oresbia has a combination of longer calyx lobes, longer pubescence, and obscurely nerved bracts, which usually serve to separate it from C. pallescens var. inverta. All three have different, though somewhat overlapping, ranges. Castilleja oresbia occasionally hybridizes with C. peckiana in Grant County, Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja cusickii includes two morphologically distinct variants. One variant has a wider inflorescence, broader bracts, and wider bract lobes. These bracts often have highly contrasting purple veins, and some populations also have a purplish wash on the bracts. These plants have a range on the western edge of typical C. cusickii, though overlapping with more typical forms in western Idaho and adjacent northeastern Oregon and southeastern Washington. The name C. lutea is available for these variants, and both consistent morphological differences and preliminary results from ongoing genetic research on the plants (D. C. Tank, pers. comm.) indicate its resurrection may be justified. A second variant is distinguished by a pale pink-purple wash on the bracts and a narrow distribution in the meadows of the Bear River Valley, bordering northwestern Utah and adjacent southeastern Idaho. Only bract coloration appears to distinguish this form from typical C. cusickii, and the distinctive bract coloration is only more or less consistent in these populations. Typical Castilleja cusickii appears to hybridize with C. gracillima in the Logan Valley, Grant County, Oregon. It is also reported by M. Ownbey (1959) to hybridize with C. miniata and C. rhexiifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Synonyms | C. lutea | |
| Name authority | Greenman: Bot. Gaz. 48: 147. (1909) | Greenman: Bot. Gaz. 25: 267. (1898) — (as Castilleia) |
| Source | FNA vol. 17, p. 635. | FNA vol. 17, p. 600. |
| Web links | ||