Castilleja nivea |
Castilleja wightii |
|
---|---|---|
snow Indian paintbrush, snow paintbrush, snowy paintbrush |
Wight's Indian paintbrush, Wight's paintbrush |
|
Habit | Herbs, perennial, 0.5–1.6 dm; from a woody caudex; with a taproot. | Herbs or subshrubs, perennial, 3–8 dm; from a woody caudex; with woody, branching roots. |
Stems | few to several, erect to ascending, decumbent at base, unbranched except for small, leafy axillary shoots, hairs weakly spreading to appressed, ± matted, especially distally on stem, fairly short and sparse proximally, longer and denser distally, soft, eglandular, becoming woolly, often obscuring surface. |
few to many, ascending to erect, much-branched, usually with prominent leafy axillary shoots, hairs spreading, long, soft, mixed with shorter, stipitate-glandular ones. |
Leaves | gray with hairs, surface green to purple, linear to narrowly lanceolate, 1–3.8 cm, not fleshy, margins plane, sometimes ± wavy, involute, 0–3-lobed, apex acute; lobes ascending-spreading, linear to narrowly lanceolate, apex acute to obtuse. |
green-tinged or ± purple, sometimes yellow-green tinged, densely crowded, linear, lanceolate, oblong, elliptic, or ovate, 1–7 cm, thickened, not usually fleshy, margins plane to wavy, involute or flat, 0–3(–5)-lobed, apex broadly rounded to obtuse or acute; lobes spreading-ascending, linear, narrowly lanceolate, triangular, or rounded, apex acute, obtuse, or rounded. |
Inflorescences | 2.5–6 × 1–2.5 cm; bracts greenish to pale yellow-green or very pale, dull purplish throughout, lanceolate to oblong, (0–)3(–5)-lobed; lobes usually ascending, linear, medium length to long, arising near mid length, apex acute. |
3–22 × 1.5–3.5 cm; bracts proximally green to deep purple, rarely light tan, distally red, scarlet, rose, red-orange, or yellow, sometimes orange, dull brownish orange, pale pinkish tan, yellow aging white, yellow aging pink, red with pink apices, magenta, or white, sometimes with a yellow to deep purple medial band, lanceolate, oblong, or narrowly ovate, (0–)3–5(–7+)-lobed, distal margins of central lobe and sometimes also side lobes with multiple shallow teeth, proximal often wavy-margined; lobes erect or ascending, linear to lanceolate or oblanceolate, short, arising in distal 2/3, apex obtuse to acute, central lobe often rounded to truncate. |
Pedicels | 0 mm or nearly so. |
|
Corollas | straight, 18–25 mm; tube 3.5–5.5 mm; subequal to calyx, or beak and sometimes abaxial lip exserted; beak adaxially yellow, 6–8 mm, hairs moderately long, matted on midline, very short-glandular on sides; abaxial lip green, inconspicuous, slightly pouched, 3.5–5.5 mm, 60–90% as long as beak; teeth erect, white or yellow, 0.5–3 mm. |
slightly curved proximally, 20–30 mm; tube 10–12 mm; distal portion of beak exserted, abaxial lip included; beak adaxially green to yellow, 11.5–18 mm; abaxial lip deep green, sometimes to very deep purple, reduced, included, usually not visible through front cleft, 1–2 mm, 15–25% as long as beak; teeth erect, green to sometimes pink, 0.5–1.5 mm. |
Calyces | yellow, color mostly obscured by whitish hairs, 15–22 mm; abaxial, adaxial, and lateral clefts (5.5–)7–12 mm, 35–55% of calyx length, often appearing shorter because matted hairs stitch proximal part of clefts shut, all 4 clefts subequal; lobes broadly linear, apex acute. |
colored as bracts, 15–28 mm; abaxial and adaxial clefts 8–11 mm, 33–50% of calyx length, deeper than laterals, lateral 2–5.5 mm, 5–30% of calyx length; lobes lanceolate to oblong to broadly triangular, apex acute, obtuse, or rounded. |
2n | = 24. |
= 24, 48. |
Castilleja nivea |
Castilleja wightii |
|
Phenology | Flowering Jun–Aug. | Flowering Mar–Aug(–Nov). |
Habitat | Gravelly slopes and flats, turf and fellfields, mostly alpine. | Coastal scrub, damp thickets, stream banks, sea bluffs, canyon slopes, roadsides. |
Elevation | 1700–3600 m. (5600–11800 ft.) | 0–700 m. (0–2300 ft.) |
Distribution |
MT; WY
|
CA
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Discussion | Castilleja nivea is endemic to alpine habitats in the mountains of northwestern Wyoming and adjacent Montana. It forms occasional hybrids with C. pulchella, which often shares its habitat, as on the Beartooth Plateau in northwestern Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja wightii is found along or near the central and northern coast of California. Historical collections of C. wightii from Curry County, Oregon, are referable to other species, but it should be sought in the area. Reports from the south-central coast of California are referable to other species, particularly C. affinis. Castilleja wightii appears to intergrade with C. latifolia and perhaps C. affinis south of San Francisco. Despite much attention from botanists, the perennial paintbrushes along the coast between Monterey and San Francisco can be perplexing and difficult to identify. This situation is likely the result of introgression, but this complex is in need of meticulous genetic and morphological analysis. North of San Francisco, C. wightii is straightforward to recognize, with its abundantly stipitate-glandular stems and leaves. In addition, the leaves are often crowded on the stems, which bear axillary shoots. Most populations have either red or yellow bracts, with only occasional individual plants of the other color. Yellow populations are found primarily in Marin and southern Sonoma counties southward and are gradually replaced by red populations from northern Sonoma County northward. Mixed color populations occur in a few places, especially in San Mateo County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 634. | FNA vol. 17, p. 664. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. wightii subsp. rubra | |
Name authority | Pennell & Ownbey: Notul. Nat. Acad. Nat. Sci. Philadelphia 227: 2. (1950) | Elmer: Bot. Gaz. 41: 322. (1906) — (as Castilleia) |
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