Castilleja nivea |
Castilleja miniata |
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snow Indian paintbrush, snow paintbrush, snowy paintbrush |
common paintbrush, common red paintbrush, giant red Indian paintbrush, great red paintbrush, scarlet Indian paintbrush, scarlet or common or giant red paintbrush, scarlet paintbrush |
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Habit | Herbs, perennial, 0.5–1.6 dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.2–8(–10) dm; from a woody caudex (or slender rooting rhizomes in var. dixonii, var. miniata); with a taproot or with slender, branched roots from a rhizome. | ||||||||||||
Stems | few to several, erect to ascending, decumbent at base, unbranched except for small, leafy axillary shoots, hairs weakly spreading to appressed, ± matted, especially distally on stem, fairly short and sparse proximally, longer and denser distally, soft, eglandular, becoming woolly, often obscuring surface. |
few to many, erect to ascending, rarely proximally decumbent or creeping and rooting at nodes, usually branched, glabrous, glabrate, or hairy, hairs spreading to ± retrorse, short to long, soft to stiff, rarely stipitate-glandular. |
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Leaves | gray with hairs, surface green to purple, linear to narrowly lanceolate, 1–3.8 cm, not fleshy, margins plane, sometimes ± wavy, involute, 0–3-lobed, apex acute; lobes ascending-spreading, linear to narrowly lanceolate, apex acute to obtuse. |
green to purple, linear to lanceolate, narrowly elliptic, narrowly oblong, or ovate, (1.5–)3–8(–9.5) cm, thin and not fleshy or slightly to moderately thickened and slightly fleshy, margins plane, rarely wavy, flat to involute, whole leaf sometimes recurved downward, 0(–5)-lobed, apex acute to obtuse, sometimes rounded; lobes ascending-spreading, narrowly lanceolate, apex acute. |
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Inflorescences | 2.5–6 × 1–2.5 cm; bracts greenish to pale yellow-green or very pale, dull purplish throughout, lanceolate to oblong, (0–)3(–5)-lobed; lobes usually ascending, linear, medium length to long, arising near mid length, apex acute. |
3–15(–22) × 1.5–5.5 cm, often bearing a thin, white, powdery exudate, especially on bract surfaces; bracts greenish, scarlet, red, red-orange, or pale orange throughout, sometimes pink, magenta, pink-purple, yellow, greenish yellow, white, or salmon throughout, or proximally greenish, distally colored as above, lanceolate to oblong-ovate, 0–5(–7)-lobed, central lobes sometimes distally apiculate; lobes erect, linear to lanceolate, oblong, or oblanceolate, short or medium length, arising near or above mid length, central lobe apex obtuse, rounded, or truncate, lateral ones rounded to acute or acuminate. |
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Pedicels | 0–5 mm. |
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Corollas | straight, 18–25 mm; tube 3.5–5.5 mm; subequal to calyx, or beak and sometimes abaxial lip exserted; beak adaxially yellow, 6–8 mm, hairs moderately long, matted on midline, very short-glandular on sides; abaxial lip green, inconspicuous, slightly pouched, 3.5–5.5 mm, 60–90% as long as beak; teeth erect, white or yellow, 0.5–3 mm. |
slightly curved, 20–48 mm; tube 12–26 mm; abaxial lip usually not exserted, though often visible in front calyx cleft, beak partially to fully exserted; beak adaxially green to yellow-green or whitish, (9–)14–25 mm; abaxial lip incurved or ascending, deep green or green, sometimes deep purple or yellowish, reduced, not inflated, visible in front cleft, 0.5–3.5 mm, 5–20% as long as beak (to ca. 33% as long as beak in some populations of var. miniata); teeth incurved or erect, green or white, 0.7–1.5 mm. |
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Calyces | yellow, color mostly obscured by whitish hairs, 15–22 mm; abaxial, adaxial, and lateral clefts (5.5–)7–12 mm, 35–55% of calyx length, often appearing shorter because matted hairs stitch proximal part of clefts shut, all 4 clefts subequal; lobes broadly linear, apex acute. |
colored as bracts, 15–38 mm; abaxial and adaxial clefts 4–24 mm, 35–70% of calyx length, deeper than laterals, lateral (1–)3–8(–12) mm, 5–30% of calyx length; lobes linear or narrowly lanceolate to narrowly triangular, apex acute to acuminate or obtuse. |
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2n | = 24. |
= 24, 48, 72, 96, 120, 144. |
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Castilleja nivea |
Castilleja miniata |
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Phenology | Flowering Jun–Aug. | |||||||||||||
Habitat | Gravelly slopes and flats, turf and fellfields, mostly alpine. | |||||||||||||
Elevation | 1700–3600 m. (5600–11800 ft.) | |||||||||||||
Distribution |
MT; WY
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AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; ON; SK; YT; nw Mexico
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Discussion | Castilleja nivea is endemic to alpine habitats in the mountains of northwestern Wyoming and adjacent Montana. It forms occasional hybrids with C. pulchella, which often shares its habitat, as on the Beartooth Plateau in northwestern Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Castilleja miniata is widely recognized as the common scarlet paintbrush. It is highly variable and has five levels of polyploidy. Nonetheless, it remains fairly well defined morphologically across its wide range. Native Americans use it medicinally. A probable hybrid with C. septentrionalis from southern Nevada was named C. ×porterae Cockerell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 634. | FNA vol. 17, p. 627. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Name authority | Pennell & Ownbey: Notul. Nat. Acad. Nat. Sci. Philadelphia 227: 2. (1950) | Douglas ex Hooker: Fl. Bor.-Amer. 2: 106. (1838) | ||||||||||||
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