Castilleja nivea
Castilleja linariifolia
snow Indian paintbrush, snow paintbrush, snowy paintbrush
desert paintbrush, desert paintbursh, linaria-leaf Indian paintbrush, narrow leaf paintbrush, narrow-leaf Indian paintbrush, Wyoming Indian paintbrush, Wyoming or narrow-leaf or long-leaf paintbrush, Wyoming paintbrush
few to several, erect to ascending, decumbent at base, unbranched except for small, leafy axillary shoots, hairs weakly spreading to appressed, ± matted, especially distally on stem, fairly short and sparse proximally, longer and denser distally, soft, eglandular, becoming woolly, often obscuring surface.
few to many, ascending to erect, much-branched, glabrous proximally, hairy distally, sometimes glabrous or hairy throughout, hairs sparse, sometimes dense, spreading to slightly retrorse, short to long, ± stiff, eglandular.
gray with hairs, surface green to purple, linear to narrowly lanceolate, 1–3.8 cm, not fleshy, margins plane, sometimes ± wavy, involute, 0–3-lobed, apex acute;
lobes ascending-spreading, linear to narrowly lanceolate, apex acute to obtuse.
± yellow to gray-green, sometimes becoming ± purple, linear to narrowly lanceolate, (1–)2–10 cm, not fleshy, rarely ± thickened and fleshy, margins plane, involute, 0–3(–5)-lobed, apex acute or acuminate;
lobes spreading, linear, apex acute to acuminate.
2.5–6 × 1–2.5 cm;
bracts greenish to pale yellow-green or very pale, dull purplish throughout, lanceolate to oblong, (0–)3(–5)-lobed;
lobes usually ascending, linear, medium length to long, arising near mid length, apex acute.
4–20 × 2–7.5 cm;
bracts red to red-orange throughout, sometimes pale green, yellow, magenta, pink-purple, or white throughout, or proximally pale greenish to straw colored, distally colored as above, lanceolate to narrowly lanceolate or narrowly oblong, 3(–5)-lobed;
lobes spreading to ascending, narrowly lanceolate to linear, shorter than central lobe, arising in proximal 1/3, apex obtuse to acuminate.
straight, 18–25 mm;
tube 3.5–5.5 mm; subequal to calyx, or beak and sometimes abaxial lip exserted;
beak adaxially yellow, 6–8 mm, hairs moderately long, matted on midline, very short-glandular on sides;
abaxial lip green, inconspicuous, slightly pouched, 3.5–5.5 mm, 60–90% as long as beak;
teeth erect, white or yellow, 0.5–3 mm.
often slightly curved, 25–45 mm;
tube 11–22(–25) mm;
beak exserted, longer than calyx lobes, usually projecting through abaxial cleft;
beak adaxially yellow-green or green, 9–21(–24) mm;
abaxial lip deep green, reduced, often visible in exserted sideways through abaxial calyx cleft, 0.5–3 mm, 10–15% as long as beak;
teeth incurved, green or whitish, 5–2(–3) mm.
yellow, color mostly obscured by whitish hairs, 15–22 mm;
abaxial, adaxial, and lateral clefts (5.5–)7–12 mm, 35–55% of calyx length, often appearing shorter because matted hairs stitch proximal part of clefts shut, all 4 clefts subequal;
lobes broadly linear, apex acute.
proximally greenish, whitish, or yellowish, distally colored as bracts, 18–30(–35) mm;
abaxial clefts 10–20(–22) mm, adaxial 2–6(–12) mm, abaxial ca. 70% of calyx length, adaxial ca. 20–25% of calyx length, deeper than laterals, lateral 1.5–5(–6) mm, 12–17% of calyx length;
lobes curved slightly toward adaxial side, narrowly oblong to narrowly lanceolate, apex acute.
= 24.
= 24, 48.
Castilleja nivea
Castilleja linariifolia
Castilleja nivea is endemic to alpine habitats in the mountains of northwestern Wyoming and adjacent Montana. It forms occasional hybrids with C. pulchella, which often shares its habitat, as on the Beartooth Plateau in northwestern Wyoming.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Castilleja linariifolia is widespread in the western United States and is important to Native Americans as a source of dyes and for medicinal and ceremonial purposes (D. E. Moerman 1998). It is the state flower of Wyoming. Castilleja linariifolia is closely associated with, and undoubtedly parasitic on, the roots of sagebrush, Artemisia tridentata. The bracts are usually red to red-orange, with uncommon yellow-bracted variants. On the eastern slope of the Sierra Nevada in California, most populations have pink-purple to magenta bracts, and at least one population has almost pure white bracts. Plants associated with hot springs in the Ash Meadows area of Nye County, Nevada, are unusually tall and have somewhat fleshy leaves. Castilleja linariifolia stems are usually glabrous, but pubescent stems are sporadic across its range. However, in the Mt. Charleston area of the Spring Mountains, in southern Nevada, most plants are pubescent, ranging from an inconspicuous layer to a fairly dense and obvious indument. These plants have been called forma omnipubescens Pennell. Elsewhere, especially in central and northern Arizona, individuals with short-pubescent stems are intermingled with more typical plants. In addition, late-blooming forms associated with hot spring sites in the eastern Mojave Desert are particularly thick-stemmed and vigorous. Hybrids between C. linariifolia and C. scabrida are known from Garfield County, Utah. Castilleja linariifolia is parapatric with the similar C. wootonii in central New Mexico. Castilleja linariifolia is reported to hybridize with C. chromosa, C. flava, and C. miniata. A possible hybrid with C. septentrionalis was named C. ×cognata Greene.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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