Castilleja montigena |
Castilleja uliginosa |
|
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Heckard's Indian paintbrush, Heckard's paintbrush |
Pitkin Marsh Indian paintbrush, Pitkin Marsh paintbrush |
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Habit | Herbs or subshrubs, perennial, 1.5–4.5 dm; from a woody caudex; with a taproot. | Herbs, perennial, 3–5 dm; from a woody caudex; rhizomatous. |
Stems | few to several, decumbent to erect, sometimes leaning, unbranched or often much-branched distally, with a few short, leafy axillary shoots, hairs spreading, short, soft, stipitate-glandular, mixed with long-spreading, eglandular ones. |
several, decumbent proximally and sometimes becoming weakly rhizomatous, becoming ascending to erect, unbranched or often branched proximally, hairs spreading, short and long, soft, mixed eglandular and stipitate-glandular. |
Leaves | gray-green, sometimes green, lanceolate-linear to narrowly lanceolate, 1–6.5 cm, not fleshy, margins plane, sometimes wavy, flat to involute, 0(–3)-lobed, apex acuminate; lobes spreading-ascending, linear to narrowly lanceolate, apex acute. |
green, lanceolate or broadly lanceolate, 3–5 cm, much reduced distally, not fleshy, margins plane, sometimes ± wavy, flat, 0-lobed, apex acute to rounded. |
Inflorescences | 3–30 × 3–4 cm; bracts proximally green to dark purplish, distally red to crimson, sometimes pale salmon, linear-lanceolate to broadly lanceolate, 3–5-lobed; lobes spreading, linear, long, arising below mid length, apex acute, sometimes obtuse. |
10–30 × 3–4 cm; bracts pale yellow to cream throughout, or proximally pale greenish near base, distally pale yellow to cream, broadly lanceolate, 3-lobed; lobes ascending, narrowly lanceolate, long, arising near mid length, apex narrowly acute or acuminate. |
Corollas | straight or slightly curved, 20–40 mm; tube 15–23 mm; abaxial lip exserted to included, beak much exserted; beak adaxially yellow-green to reddish, 9–18 mm; abaxial lip green, reduced, 0.5–1.5 mm, 5–20% as long as beak; teeth incurved, green, (0–)0.5–1.5 mm. |
slightly curved, 22–30 mm; tube 11–15 mm; beak partly exserted, adaxially whitish, yellowish, or greenish, 10–15 mm; abaxial lip green, small, protuberant, 3 mm, ca. 20% as long as beak; teeth erect to incurved, green, to 1 mm. |
Calyces | colored as bracts, 15–20 mm; abaxial clefts 3.4–6.2 mm, adaxial 4.5–9 mm, clefts 25–33% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–10% of calyx length; lobes narrowly triangular, often slightly unequal, apex acute. |
colored as bracts, 20–25 mm; abaxial and adaxial clefts 7–14 mm, ca. 50% of calyx length, deeper than laterals, lateral 5–6 mm, ca. 33% of calyx length; lobes linear, apex acute, ciliate. |
2n | = 48, 72. |
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Castilleja montigena |
Castilleja uliginosa |
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Phenology | Flowering May–Aug. | Flowering May–Jun. |
Habitat | Dry rocky slopes, ledges, open conifer forests, thickets, washes. | Margins of wet meadows, marshes, and wet thickets. |
Elevation | 1900–2900 m. (6200–9500 ft.) | 40–60 m. (100–200 ft.) |
Distribution |
CA
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CA |
Discussion | Castilleja montigena is endemic to the northeastern portion of the San Bernardino Mountains of southern California. In the field, this species is consistently and relatively easily distinguished from nearby populations of C. martini var. martini, which it essentially replaces in the northeastern portion of the San Bernardino Mountains. It is apparently of allopolyploid hybrid origin between C. martini var. martini and C. chromosa, which approaches its range from the adjacent Mojave Desert. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja uliginosa is endemic to Pitkin Marsh in Sonoma County. It differs from C. miniata by its uniformly pale yellow inflorescences and pubescent stems, as well as by its disjunct, low-elevation habitat. Much of its available habitat was destroyed by development, and it is apparently now extirpated from the wild. It survives in the form of tissue clones from the last wild plant, backcrossed with C. miniata by L. R. Heckard in the 1980s and maintained since then in the greenhouses at the University of California at Berkeley. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 632. | FNA vol. 17, p. 662. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Name authority | Heckard: Syst. Bot. 5: 83, fig. 17 [center]. (1980) | Eastwood: Leafl. W. Bot. 3: 117. (1942) |
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