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Heckard's Indian paintbrush, Heckard's paintbrush

yoho paintbrush

Habit Herbs or subshrubs, perennial, 1.5–4.5 dm; from a woody caudex; with a taproot. Herbs, perennial, (1.5–)2–4 dm; from a woody caudex; with branched, woody roots.
Stems

few to several, decumbent to erect, sometimes leaning, unbranched or often much-branched distally, with a few short, leafy axillary shoots, hairs spreading, short, soft, stipitate-glandular, mixed with long-spreading, eglandular ones.

few or several, ascending to erect, branched or unbranched, shiny proximally, glabrous proximally, hairy distally, hairs spreading, long, soft, eglandular, sometimes sparsely glandular.

Leaves

gray-green, sometimes green, lanceolate-linear to narrowly lanceolate, 1–6.5 cm, not fleshy, margins plane, sometimes wavy, flat to involute, 0(–3)-lobed, apex acuminate;

lobes spreading-ascending, linear to narrowly lanceolate, apex acute.

deep purple to green, linear-lanceolate to narrowly lanceolate, rarely broadly lanceolate, 1.5–4.5 cm, not fleshy, margins plane, flat or slightly involute, 0(–3)-lobed, apex usually acuminate or acute;

lobes upright or ascending, lanceolate, apex acute.

Inflorescences

3–30 × 3–4 cm;

bracts proximally green to dark purplish, distally red to crimson, sometimes pale salmon, linear-lanceolate to broadly lanceolate, 3–5-lobed;

lobes spreading, linear, long, arising below mid length, apex acute, sometimes obtuse.

2–12 × 2–6 cm;

bracts deep reddish or crimson, rarely magenta or dull orange, or proximally greenish near base, distally colored as above, oblong to broadly lanceolate, often spreading from base and exposing calyces, 0–3(–5)-lobed;

lobes ascending to erect, linear, short to medium length, arising near or above mid length, apex acute to obtuse.

Corollas

straight or slightly curved, 20–40 mm;

tube 15–23 mm;

abaxial lip exserted to included, beak much exserted;

beak adaxially yellow-green to reddish, 9–18 mm;

abaxial lip green, reduced, 0.5–1.5 mm, 5–20% as long as beak;

teeth incurved, green, (0–)0.5–1.5 mm.

straight to slightly curved, 20–37 mm;

tube 12–21 mm;

abaxial lip sometimes exserted, beak exserted;

beak adaxially green, 10–16 mm;

abaxial lip green, sometimes yellow, usually reduced, protruding through abaxial cleft, sometimes a little pouched, 3-lobed, 2–6 mm, 17–45% as long as beak;

teeth erect, green, whitish, or yellow, 1–1.5 mm.

Calyces

colored as bracts, 15–20 mm;

abaxial clefts 3.4–6.2 mm, adaxial 4.5–9 mm, clefts 25–33% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–10% of calyx length;

lobes narrowly triangular, often slightly unequal, apex acute.

colored as bracts, 18–28 mm;

abaxial clefts 5–6 mm, adaxial 8–15 mm, clefts 28–50% of calyx length, deeper than laterals, lateral 1.5–6 mm, 5–25% of calyx length;

lobes long-triangular, apex acute.

2n

= 48, 72.

= 48.

Castilleja montigena

Castilleja purpurascens

Phenology Flowering May–Aug. Flowering (May–)Jun–Jul.
Habitat Dry rocky slopes, ledges, open conifer forests, thickets, washes. Gravel river flats, moist thickets.
Elevation 1900–2900 m. (6200–9500 ft.) 1200–2200 m. (3900–7200 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AB; BC
Discussion

Castilleja montigena is endemic to the northeastern portion of the San Bernardino Mountains of southern California. In the field, this species is consistently and relatively easily distinguished from nearby populations of C. martini var. martini, which it essentially replaces in the northeastern portion of the San Bernardino Mountains. It is apparently of allopolyploid hybrid origin between C. martini var. martini and C. chromosa, which approaches its range from the adjacent Mojave Desert.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Castilleja purpurascens is a species of gravelly flood plains and riverbanks at moderate elevations in the vicinity of the Kicking Horse River in Yoho National Park, British Columbia, and in immediately adjacent Alberta. While this tetraploid species may be derived from hybridization between C. miniata and C. rhexiifolia, its combination of traits is unique, and the species forms morphologically consistent populations limited to a habitat not particularly favored by either putative parent species. Very occasional hybrids are known with both C. lutescens and C. miniata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 632. FNA vol. 17, p. 649.
Parent taxa Orobanchaceae > Castilleja Orobanchaceae > Castilleja
Sibling taxa
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. disticha, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lanata, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. martini, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurascens, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. wootonii, C. xanthotricha
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. disticha, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lanata, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. martini, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. montigena, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. wootonii, C. xanthotricha
Name authority Heckard: Syst. Bot. 5: 83, fig. 17 [center]. (1980) Greenman: Bot. Gaz. 42: 146. (1906)
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