Castilleja montigena |
Castilleja hyperborea |
|
---|---|---|
Heckard's Indian paintbrush, Heckard's paintbrush |
northern Indian paintbrush, northern paintbrush |
|
Habit | Herbs or subshrubs, perennial, 1.5–4.5 dm; from a woody caudex; with a taproot. | Herbs, perennial, 0.7–2.6 dm; from a many-headed, short, woody caudex; with a taproot. |
Stems | few to several, decumbent to erect, sometimes leaning, unbranched or often much-branched distally, with a few short, leafy axillary shoots, hairs spreading, short, soft, stipitate-glandular, mixed with long-spreading, eglandular ones. |
few, ascending, unbranched, sometimes branched, glabrate to sparsely hairy proximally, becoming densely hairy distally, proximal hairs appressed, white, short, soft, eglandular, distal hairs widely spreading to erect, white or yellowish, long to very long, soft, eglandular. |
Leaves | gray-green, sometimes green, lanceolate-linear to narrowly lanceolate, 1–6.5 cm, not fleshy, margins plane, sometimes wavy, flat to involute, 0(–3)-lobed, apex acuminate; lobes spreading-ascending, linear to narrowly lanceolate, apex acute. |
green to sometimes red-brown or dull purplish, linear to linear-lanceolate, 1.5–4 cm, not fleshy, margins plane, sometimes ± wavy, involute, (0–)3(–5)-lobed, apex acute to acuminate; lobes linear or linear-lanceolate, laterals ± divaricate, often abruptly up-curved from plane of main leaf blade, proximals usually 33–50% as long as leaf, apex acuminate to acute. |
Inflorescences | 3–30 × 3–4 cm; bracts proximally green to dark purplish, distally red to crimson, sometimes pale salmon, linear-lanceolate to broadly lanceolate, 3–5-lobed; lobes spreading, linear, long, arising below mid length, apex acute, sometimes obtuse. |
2–11 × 1–2.5 cm; bracts greenish or pale yellow, rarely purplish or deep red, often with dull purplish veins on proximal bracts, or proximally greenish or pale yellow, distally whitish, yellow, or orange-yellow, proximal sometimes linear-lanceolate, distal broadly lanceolate to oblong or obovate, (0–)3–5(–7)-lobed; lobes ascending to erect, linear to lanceolate, short to medium length, arising above mid length, proximal bract apex acute, distal obtuse, rounded, or truncate. |
Corollas | straight or slightly curved, 20–40 mm; tube 15–23 mm; abaxial lip exserted to included, beak much exserted; beak adaxially yellow-green to reddish, 9–18 mm; abaxial lip green, reduced, 0.5–1.5 mm, 5–20% as long as beak; teeth incurved, green, (0–)0.5–1.5 mm. |
slightly curved, 10–22 mm; tube 8–14 mm; teeth of abaxial lip sometimes slightly exserted, beak exserted or subequal; beak adaxially green to pale yellow, 5–8 mm; abaxial lip deep yellow or green, reduced, forming a small pouch visible through front cleft, 3–5 mm, 50–75% as long as beak; teeth ascending to erect, white or yellow, 1 mm. |
Calyces | colored as bracts, 15–20 mm; abaxial clefts 3.4–6.2 mm, adaxial 4.5–9 mm, clefts 25–33% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–10% of calyx length; lobes narrowly triangular, often slightly unequal, apex acute. |
colored as bracts, proximally often paler, usually yellow throughout, 10–20 mm; abaxial and adaxial clefts 7–14 mm, 40–50% of calyx length, deeper than laterals, lateral 0.5–1.5 mm, 5–15% of calyx length; lobes triangular, apex acute to rounded. |
2n | = 48, 72. |
= 24. |
Castilleja montigena |
Castilleja hyperborea |
|
Phenology | Flowering May–Aug. | Flowering May–Aug(–Sep). |
Habitat | Dry rocky slopes, ledges, open conifer forests, thickets, washes. | Dry to mesic rocky tundra, slopes, ridges, barrens and meadows, openings in boreal forests, arctic to alpine. |
Elevation | 1900–2900 m. (6200–9500 ft.) | 0–2200 m. (0–7200 ft.) |
Distribution |
CA
|
AK; BC; NT; YT; e Asia (Russian Far East) |
Discussion | Castilleja montigena is endemic to the northeastern portion of the San Bernardino Mountains of southern California. In the field, this species is consistently and relatively easily distinguished from nearby populations of C. martini var. martini, which it essentially replaces in the northeastern portion of the San Bernardino Mountains. It is apparently of allopolyploid hybrid origin between C. martini var. martini and C. chromosa, which approaches its range from the adjacent Mojave Desert. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja hyperborea is widespread across boreal, alpine, and arctic habitats in western Canada and Alaska, extending into the Russian Far East. A rare form with light purple bracts was named as C. kuschei. Another form with particularly long hairs found in the Kluane Lake region of the southern Yukon was described as C. villosissima and is usually found on calcareous substrates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 632. | FNA vol. 17, p. 614. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. kuschei, C. villosissima | |
Name authority | Heckard: Syst. Bot. 5: 83, fig. 17 [center]. (1980) | Pennell: Proc. Acad. Nat. Sci. Philadelphia 86: 532. (1934) |
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