Castilleja montigena
Castilleja hololeuca
Heckard's Indian paintbrush, Heckard's paintbrush
Channel Islands paintbrush, island paintbrush, island white-felted paintbrush, white-felted paintbrush
few to several, decumbent to erect, sometimes leaning, unbranched or often much-branched distally, with a few short, leafy axillary shoots, hairs spreading, short, soft, stipitate-glandular, mixed with long-spreading, eglandular ones.
many, erect to spreading, much-branched with many short axillary shoots, proximal stems reaching 1+ cm wide, white-felted, hairs dense, spreading to ± appressed, intertwined, long, soft, slightly branched, obscuring surface.
gray-green, sometimes green, lanceolate-linear to narrowly lanceolate, 1–6.5 cm, not fleshy, margins plane, sometimes wavy, flat to involute, 0(–3)-lobed, apex acuminate;
lobes spreading-ascending, linear to narrowly lanceolate, apex acute.
ash gray, leaves of previous year persisting on proximal stem, linear, older leaves sometimes broadly linear, 1–5 cm, not fleshy, margins plane, involute, 0-lobed, apex obtuse;
lobes ascending, linear-lanceolate, apex obtuse to rounded, sometimes acute.
3–30 × 3–4 cm;
bracts proximally green to dark purplish, distally red to crimson, sometimes pale salmon, linear-lanceolate to broadly lanceolate, 3–5-lobed;
lobes spreading, linear, long, arising below mid length, apex acute, sometimes obtuse.
3–16 × 2–4 cm;
bracts pale gray-green throughout, or proximally pale gray-green, distally red or yellow, proximal linear, distal broader, often oblanceolate to obovate, (0–)3-lobed;
lobes spreading, linear to oblong, short to long, arising below mid length, apex rounded to truncate or acute.
straight or slightly curved, 20–40 mm;
tube 15–23 mm;
abaxial lip exserted to included, beak much exserted;
beak adaxially yellow-green to reddish, 9–18 mm;
abaxial lip green, reduced, 0.5–1.5 mm, 5–20% as long as beak;
teeth incurved, green, (0–)0.5–1.5 mm.
slightly curved in proximal 1/3, 14–26(–31) mm;
tube 9–17 mm;
abaxial lip not exserted, beak exserted;
beak adaxially green to yellowish, 11–14 mm;
abaxial lip ascending, deep green, reduced, 2–3 mm, less than 33% as long as beak;
teeth reduced to apiculations, deep green, 1–1.5 mm.
colored as bracts, 15–20 mm;
abaxial clefts 3.4–6.2 mm, adaxial 4.5–9 mm, clefts 25–33% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–10% of calyx length;
lobes narrowly triangular, often slightly unequal, apex acute.
colored as bracts, with conspicuous whitish veins, 14–18(–20) mm;
abaxial and adaxial clefts 5–12 mm, 30–67% of calyx length, deeper than laterals, lateral 0 mm or nearly so, ca. 0% of calyx length;
lobes linear-lanceolate, apex rounded, truncate, or emarginate.
= 48, 72.
Castilleja montigena
Castilleja hololeuca
Castilleja montigena is endemic to the northeastern portion of the San Bernardino Mountains of southern California. In the field, this species is consistently and relatively easily distinguished from nearby populations of C. martini var. martini, which it essentially replaces in the northeastern portion of the San Bernardino Mountains. It is apparently of allopolyploid hybrid origin between C. martini var. martini and C. chromosa, which approaches its range from the adjacent Mojave Desert.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Castilleja hololeuca is endemic to the four major islands of the northern Channel Islands. It is similar to C. lanata, a mostly Mexican species that reaches Arizona, New Mexico, and southwest Texas. However, C. hololeuca appears to be more closely related to C. foliolosa, a species of the California mainland. These three species, and C. galehintoniae G. L. Nesom of Nuevo León, all share a distinctive, thick indument of pale, matted, and often branched hairs on the stems and leaves. Large individuals of C. hololeuca form thick, woody trunks. On Anacapa Island, closest to the mainland, red-flowered forms of C. hololeuca predominate. On Santa Cruz Island, to the west, both red- and yellow-bracted forms are fairly common, usually in separate populations. Further offshore, on San Miguel and Santa Rosa islands, most plants are yellow to, occasionally, peach in coloration. Reports of this species from Santa Barbara and Santa Catalina Islands are referable to other species, mostly C. foliolosa. Populations of C. hololeuca historically declined from grazing by introduced game and livestock species. The plants are recovering well, following removal of the grazing animals, and are now common in many areas, especially on San Miguel Island.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
CA
WildflowerSearch
iNaturalist
USDA Plants Database
