Castilleja miniata |
Castilleja nana |
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common paintbrush, common red paintbrush, giant red Indian paintbrush, great red paintbrush, scarlet Indian paintbrush, scarlet or common or giant red paintbrush, scarlet paintbrush |
alpine paintbrush, dwarf alpine Indian paintbrush, dwarf alpine paintbrush, dwarf paintbrush |
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Habit | Herbs, perennial, 1.2–8(–10) dm; from a woody caudex (or slender rooting rhizomes in var. dixonii, var. miniata); with a taproot or with slender, branched roots from a rhizome. | Herbs, perennial, 0.4–1.7 dm; from a woody caudex; with a taproot. | ||||||||||||
Stems | few to many, erect to ascending, rarely proximally decumbent or creeping and rooting at nodes, usually branched, glabrous, glabrate, or hairy, hairs spreading to ± retrorse, short to long, soft to stiff, rarely stipitate-glandular. |
few to several, ascending to decumbent-based, unbranched, hairs spreading, long, soft to stiff, mixed with shorter stipitate-glandular ones. |
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Leaves | green to purple, linear to lanceolate, narrowly elliptic, narrowly oblong, or ovate, (1.5–)3–8(–9.5) cm, thin and not fleshy or slightly to moderately thickened and slightly fleshy, margins plane, rarely wavy, flat to involute, whole leaf sometimes recurved downward, 0(–5)-lobed, apex acute to obtuse, sometimes rounded; lobes ascending-spreading, narrowly lanceolate, apex acute. |
green to deep purple, linear to narrowly lanceolate, (0.5–)1–2.5(–3.1) cm, not fleshy, margins plane, flat to slightly involute, 3(–7)-lobed, apex acuminate; lobes ascending-erect, linear to filiform, apex acute to obtuse. |
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Inflorescences | 3–15(–22) × 1.5–5.5 cm, often bearing a thin, white, powdery exudate, especially on bract surfaces; bracts greenish, scarlet, red, red-orange, or pale orange throughout, sometimes pink, magenta, pink-purple, yellow, greenish yellow, white, or salmon throughout, or proximally greenish, distally colored as above, lanceolate to oblong-ovate, 0–5(–7)-lobed, central lobes sometimes distally apiculate; lobes erect, linear to lanceolate, oblong, or oblanceolate, short or medium length, arising near or above mid length, central lobe apex obtuse, rounded, or truncate, lateral ones rounded to acute or acuminate. |
15–20 × 1.5–3 cm; bracts greenish or deep purplish throughout, or proximally greenish or deep purplish, distally white, pink, magenta, reddish purple, purple, or pale yellow, distal coloration often limited to apices and margins, broadly lanceolate or narrowly to broadly elliptic, 0–3(–7)-lobed; lobes spreading, linear to narrowly oblanceolate, long, arising near mid length, apex acute or obtuse. |
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Pedicels | 0–5 mm. |
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Corollas | slightly curved, 20–48 mm; tube 12–26 mm; abaxial lip usually not exserted, though often visible in front calyx cleft, beak partially to fully exserted; beak adaxially green to yellow-green or whitish, (9–)14–25 mm; abaxial lip incurved or ascending, deep green or green, sometimes deep purple or yellowish, reduced, not inflated, visible in front cleft, 0.5–3.5 mm, 5–20% as long as beak (to ca. 33% as long as beak in some populations of var. miniata); teeth incurved or erect, green or white, 0.7–1.5 mm. |
straight, 10–16(–19) mm; tube 8–13 mm, with patches of blackish coloration on either side of distal portion; beak subequal to calyx or exserted, adaxially green, yellow, whitish, or pink, 3–5.5 mm, scarcely exceeding abaxial lip, margins brown or burgundy, sometimes pink; abaxial lip pale yellow, white, green, or purple, inflated, pouched, 2–5 mm, 65–95% as long as beak; teeth erect to slightly spreading, white, yellow, or pink, 0.5–2.1 mm. |
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Calyces | colored as bracts, 15–38 mm; abaxial and adaxial clefts 4–24 mm, 35–70% of calyx length, deeper than laterals, lateral (1–)3–8(–12) mm, 5–30% of calyx length; lobes linear or narrowly lanceolate to narrowly triangular, apex acute to acuminate or obtuse. |
green to purple, margins green, white, or pink, 10–19 mm; abaxial, adaxial, and lateral clefts 3.5–7(–10) mm, 33–55% of calyx length, all 4 clefts subequal; lobes linear, lanceolate-elliptic, or narrowly triangular, apex acute to rounded. |
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Stigmas | black. |
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2n | = 24, 48, 72, 96, 120, 144. |
= 24. |
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Castilleja miniata |
Castilleja nana |
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Phenology | Flowering Jun–Aug. | |||||||||||||
Habitat | Rocky or gravelly slopes, talus, ridges, fellfields, subalpine and alpine, often over granite. | |||||||||||||
Elevation | 1900–4300 m. (6200–14100 ft.) | |||||||||||||
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; ON; SK; YT; nw Mexico
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CA; NV; UT
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Discussion | Varieties 4 (4 in the flora). Castilleja miniata is widely recognized as the common scarlet paintbrush. It is highly variable and has five levels of polyploidy. Nonetheless, it remains fairly well defined morphologically across its wide range. Native Americans use it medicinally. A probable hybrid with C. septentrionalis from southern Nevada was named C. ×porterae Cockerell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja nana is limited to high elevations in the Sierra Nevada of California and the Great Basin ranges in central Nevada and western Utah. Plants with pink to purple inflorescences were described as C. lapidicola. Localized hybrid swarms between C. nana and C. viscidula are known from several mountain ranges in central and northern Nevada. Castilleja nana is sometimes confused with higher elevation forms of C. pilosa but can usually be distinguished from that species by the blackish patches on the sides of the corolla tube. Divergent populations in the central Sierra Nevada and adjacent White Mountains deserve further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 627. | FNA vol. 17, p. 633. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Synonyms | C. lapidicola | |||||||||||||
Name authority | Douglas ex Hooker: Fl. Bor.-Amer. 2: 106. (1838) | Eastwood: Proc. Calif. Acad. Sci., ser. 3, 2: 289. (1902) — (as Castilleia) | ||||||||||||
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