Castilleja miniata |
Castilleja lanata |
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common paintbrush, common red paintbrush, giant red Indian paintbrush, great red paintbrush, scarlet Indian paintbrush, scarlet or common or giant red paintbrush, scarlet paintbrush |
Sierra woolly Indian paintbrush, Sierra woolly paintbrush, woolly paintbrush |
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Habit | Herbs, perennial, 1.2–8(–10) dm; from a woody caudex (or slender rooting rhizomes in var. dixonii, var. miniata); with a taproot or with slender, branched roots from a rhizome. | Subshrubs or herbs, perennial, 1.8–9(–10) dm; from a woody caudex; with a thick, woody taproot or system to thick, branched roots. | ||||||||||||
Stems | few to many, erect to ascending, rarely proximally decumbent or creeping and rooting at nodes, usually branched, glabrous, glabrate, or hairy, hairs spreading to ± retrorse, short to long, soft to stiff, rarely stipitate-glandular. |
few to many, erect to ascending, unbranched, sometimes branched distally, sometimes with leafy axillary shoots, lanate to woolly, hairs dense, spreading to weakly appressed, white to yellowish, medium length, soft, moderately branched or unbranched in different populations, sometimes mixed, eglandular, rarely short stipitate-glandular, mostly obscuring surface. |
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Leaves | green to purple, linear to lanceolate, narrowly elliptic, narrowly oblong, or ovate, (1.5–)3–8(–9.5) cm, thin and not fleshy or slightly to moderately thickened and slightly fleshy, margins plane, rarely wavy, flat to involute, whole leaf sometimes recurved downward, 0(–5)-lobed, apex acute to obtuse, sometimes rounded; lobes ascending-spreading, narrowly lanceolate, apex acute. |
green to purple, often hidden by matted, white hairs, linear to narrowly oblong or narrowly lanceolate, (1–)2–7(–10) cm, not fleshy, margins plane, flat, sometimes involute, 0–3-lobed, apex rounded, sometimes acute; lobes spreading, linear or narrowly oblong, apex rounded to acute. |
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Inflorescences | 3–15(–22) × 1.5–5.5 cm, often bearing a thin, white, powdery exudate, especially on bract surfaces; bracts greenish, scarlet, red, red-orange, or pale orange throughout, sometimes pink, magenta, pink-purple, yellow, greenish yellow, white, or salmon throughout, or proximally greenish, distally colored as above, lanceolate to oblong-ovate, 0–5(–7)-lobed, central lobes sometimes distally apiculate; lobes erect, linear to lanceolate, oblong, or oblanceolate, short or medium length, arising near or above mid length, central lobe apex obtuse, rounded, or truncate, lateral ones rounded to acute or acuminate. |
2–19 × 1.5–5 cm; bracts proximally pale greenish to greenish gray, distally bright red to orange-red, sometimes pinkish, magenta, or salmon, rarely yellow, oblong to narrowly lanceolate or narrowly oblanceolate, deeply 0–3(–5)-lobed; lobes spreading, oblanceolate or linear, long, arising below mid length, distal bract apex usually rounded. |
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Pedicels | 0–5 mm. |
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Corollas | slightly curved, 20–48 mm; tube 12–26 mm; abaxial lip usually not exserted, though often visible in front calyx cleft, beak partially to fully exserted; beak adaxially green to yellow-green or whitish, (9–)14–25 mm; abaxial lip incurved or ascending, deep green or green, sometimes deep purple or yellowish, reduced, not inflated, visible in front cleft, 0.5–3.5 mm, 5–20% as long as beak (to ca. 33% as long as beak in some populations of var. miniata); teeth incurved or erect, green or white, 0.7–1.5 mm. |
curved in proximal 1/3, 23–35(–42) mm; tube 12–17 mm; beak exserted, adaxially greenish to yellowish, 11–22 mm; abaxial lip dark green, reduced, 1–4 mm, ca. 20% as long as beak; teeth incurved, reduced, green, 0.5–2.5 mm. |
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Calyces | colored as bracts, 15–38 mm; abaxial and adaxial clefts 4–24 mm, 35–70% of calyx length, deeper than laterals, lateral (1–)3–8(–12) mm, 5–30% of calyx length; lobes linear or narrowly lanceolate to narrowly triangular, apex acute to acuminate or obtuse. |
proximally whitish to green, central band whitish, distal 1/3 colored as bract lobes, sometimes lighter or deeper, 15–29 mm; abaxial and adaxial clefts 6–14 mm, ca. 50% of calyx length, deeper than laterals, lateral 0–5 mm, 0–5(–15)% of calyx length; lobes oblong to broadly triangular, apex rounded, truncate, or emarginate. |
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2n | = 24, 48, 72, 96, 120, 144. |
= 24. |
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Castilleja miniata |
Castilleja lanata |
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Phenology | Flowering year-round. | |||||||||||||
Habitat | Dry rocky slopes, ridges, canyons, flats, valleys, montane. | |||||||||||||
Elevation | 300–2300 m. (1000–7500 ft.) | |||||||||||||
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; ON; SK; YT; nw Mexico
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AZ; NM; TX; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
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Discussion | Varieties 4 (4 in the flora). Castilleja miniata is widely recognized as the common scarlet paintbrush. It is highly variable and has five levels of polyploidy. Nonetheless, it remains fairly well defined morphologically across its wide range. Native Americans use it medicinally. A probable hybrid with C. septentrionalis from southern Nevada was named C. ×porterae Cockerell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most records of Castilleja lanata are from the southern third of Arizona, New Mexico, and southwestern Texas. Plants sometimes referred to C. lanata from the Channel Islands of California are assigned here to C. hololeuca, an insular endemic. In northern Mexico, C. lanata extends from San Luis Potosí and Nuevo León westward to Sonora, with a disjunct distribution in central Baja California, where it overlaps the southern limits of a related species, C. foliolosa. Castilleja lanata inhabits a wide spectrum of soils and elevations and does not co-occur with the closely related C. galehintoniae in Nuevo León, where the latter is limited to limestone and gypsum substrates in a narrow elevational range. Hybrids of C. lanata with C. sessiliflora are known from Pecos County, Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 627. | FNA vol. 17, p. 618. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Name authority | Douglas ex Hooker: Fl. Bor.-Amer. 2: 106. (1838) | A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 118. (1859) | ||||||||||||
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