Castilleja lineata |
Castilleja applegatei |
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linear-lobed paintbrush, lineated paintbrush, marsh-meadow Indian paintbrush |
Applegate's Indian paintbrush, Applegate's paintbrush, wavy leaf paintbrush, wavy-leaf Indian paintbrush, waxy-leaf Indian paintbrush |
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Habit | Herbs, perennial, 1–4 dm; from a woody caudex; with a taproot. | Herbs, perennial, (0.8–)1–5(–6) dm; from a branched, woody caudex; with a taproot. | ||||||||||||
Stems | few to many, erect or strongly ascending, unbranched except for small, leafy axillary shoots, hairs spreading-reflexed to ± appressed, matted, long, soft, with much shorter stipitate-glandular ones, white-woolly. |
few to many, erect, ascending, or decumbent, unbranched or branched, hairs sparse to dense, spreading, long, soft to ± stiff, eglandular, mixed with shorter stipitate-glandular ones. |
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Leaves | green, linear to narrowly oblong or narrowly lanceolate, 1.3–5 cm, not fleshy, margins plane, sometimes wavy, involute, 3–7-lobed, apex acute; lobes divergent, spreading-ascending, linear, apex acute to acuminate. |
green to purplish or brown, linear to broadly lanceolate, sometimes ovate, 1–6 cm, not fleshy, margins wavy, involute, (0–)3(–5)-lobed, apex rounded or narrowly acute to acuminate; lobes spreading or ascending, lanceolate to broadly lanceolate, often short, apex acute to rounded. |
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Inflorescences | 5–22 × 1–4.5 cm; bracts green to yellow-green throughout, or proximally green to yellow-green, distally yellow to sometimes pale yellow-orange, narrowly lanceolate to narrowly oblong or broadly lanceolate, 3(–7)-lobed; lobes ascending to spreading, linear to oblong, medium length to long, arising near or below mid length, central lobe apex rounded to obtuse, lateral ones acute. |
2–12(–21 in fruit) × 1–5 cm; bracts proximally green to dull purplish brown, distally red, red-orange, or scarlet, sometimes orange, white, or yellow, rarely with a narrow yellowish band medially, lanceolate, broadly lanceolate, oblong, or lanceolate-ovate, 3–5(–7)-lobed; lobes spreading to ascending, linear, sometimes expanded near tip, long, arising from ca. mid length, central lobe apex obtuse to rounded, sometimes expanded, others acute, rounded, obtuse, or acuminate. |
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Corollas | straight or slightly curved, 14–22 mm; tube 7–14 mm; beak tip barely exserted from calyx; beak adaxially greenish, 4–7 mm; abaxial lip green to yellow, reduced, 1–4 mm, usually less than 67% as long as beak; teeth erect, white to yellow, 1–2.5 mm. |
straight or curved in proximal 1/3, 16–35(–41) mm; tube 9–22 mm; beak usually long-exserted, adaxially green, yellow-green, or yellow, 6–20 mm; abaxial lip deep green to whitish or yellow, reduced, inconspicuous, protuberant, thickened, included or exserted, 1–3 mm, 5–20% as long as beak; teeth ascending or incurved, deep green to yellow, 0.5–1 mm. |
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Calyces | colored as bracts, 15–20 mm; abaxial and adaxial clefts 5.5–8 mm, 30–50% of calyx length, ± deeper than laterals, sometimes appearing subequal in pressed specimens, lateral 5–6 mm, ca. 33% of calyx length; lobes linear to narrowly lanceolate, sometimes expanded towards apices, apex acute. |
proximally green or whitish, sometimes yellow, lobes colored as bract lobes, sometimes with a yellow band between proximal and distal portions, 13–25 mm; abaxial and adaxial clefts 4–14 mm, 33–50% of calyx length, deeper than laterals, lateral 2.5–8 mm, 12–50% of calyx length; lobes lanceolate-acuminate, narrowly oblong, or narrowly triangular, apex acuminate, acute, or obtuse. |
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2n | = 24. |
= 24, 48. |
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Castilleja lineata |
Castilleja applegatei |
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Phenology | Flowering Jun–Aug. | |||||||||||||
Habitat | Dry to moist slopes and meadows, shores, open conifer forests, montane to alpine. | |||||||||||||
Elevation | 2100–3800 m. (6900–12500 ft.) | |||||||||||||
Distribution |
AZ; CO; NM
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CA; ID; NV; OR; UT; WY
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Discussion | Castilleja lineata is restricted to the mountains of northeastern Arizona, southern Colorado, and northwestern New Mexico. It is uncommon throughout its range and is without apparent close relatives. The Navajo used C. lineata as a medicinal plant and for its sweet nectar (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Castilleja applegatei is a widespread and often common species, with complex patterns of variation and several common but inconstant color forms, especially in var. pinetorum. Castilleja disticha and C. martini, although sometimes included as subspecies within C. applegatei, are treated as species here. Both are more morphologically divergent and more easily distinguished from typical C. applegatei than are the four varieties accepted here. Variety pinetorum is the most widespread form and also occurs over a wider range of elevations than the other three varieties, which are primarily montane to subalpine and occasionally alpine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 623. | FNA vol. 17, p. 587. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Name authority | Greene: Pittonia 4: 151. (1900) — (as Castilleia) | Fernald: Erythea 6: 49. (1898) — (as Castilleia) | ||||||||||||
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