Castilleja lineariloba |
Castilleja nana |
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pale owl-clover, thin-lobed owl's clover |
alpine paintbrush, dwarf alpine Indian paintbrush, dwarf alpine paintbrush, dwarf paintbrush |
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Habit | Herbs, annual, 1.5–4.5 dm; with fibrous roots. | Herbs, perennial, 0.4–1.7 dm; from a woody caudex; with a taproot. |
Stems | solitary or few, erect, unbranched or branched, hairs spreading, short to long, ± stiff, mixed with short stipitate-glandular ones. |
few to several, ascending to decumbent-based, unbranched, hairs spreading, long, soft to stiff, mixed with shorter stipitate-glandular ones. |
Leaves | green, linear to narrowly lanceolate or narrowly oblong, 2–5.7 cm, not fleshy, margins plane, flat, 3–7(–9)-lobed, apex acuminate to acute; lobes spreading to ascending, linear to narrowly oblong, apex acuminate. |
green to deep purple, linear to narrowly lanceolate, (0.5–)1–2.5(–3.1) cm, not fleshy, margins plane, flat to slightly involute, 3(–7)-lobed, apex acuminate; lobes ascending-erect, linear to filiform, apex acute to obtuse. |
Inflorescences | 2–14 × 1–4 cm; bracts greenish throughout, or proximally greenish, distally white, cream, pale pink, or pale purple on apices, linear-lanceolate, 5–7(–9)-lobed, sometimes with secondary lobes; lobes ascending to spreading, linear to narrowly oblanceolate, long, arising all along leaf axis, apex acute to obtuse. |
15–20 × 1.5–3 cm; bracts greenish or deep purplish throughout, or proximally greenish or deep purplish, distally white, pink, magenta, reddish purple, purple, or pale yellow, distal coloration often limited to apices and margins, broadly lanceolate or narrowly to broadly elliptic, 0–3(–7)-lobed; lobes spreading, linear to narrowly oblanceolate, long, arising near mid length, apex acute or obtuse. |
Corollas | straight or slightly curved, 12–25 mm; tube 9–14 mm, expanded distally; abaxial lip sometimes slightly exserted, never hidden by slender calyx lobes, beak exserted; beak straight, adaxially white or lilac pink, 3–5.5 mm, inconspicuously puberulent; abaxial lip proximally white, distally yellow, with purple or red-brown spots, conspicuous, pouches 3, inflated, 4–6 mm, 4–5 mm wide, 2 mm deep, longer than deep, 1.5–4 mm, 90% as long as beak; teeth erect, white, usually with purple spot at base, 0.5–1 mm. |
straight, 10–16(–19) mm; tube 8–13 mm, with patches of blackish coloration on either side of distal portion; beak subequal to calyx or exserted, adaxially green, yellow, whitish, or pink, 3–5.5 mm, scarcely exceeding abaxial lip, margins brown or burgundy, sometimes pink; abaxial lip pale yellow, white, green, or purple, inflated, pouched, 2–5 mm, 65–95% as long as beak; teeth erect to slightly spreading, white, yellow, or pink, 0.5–2.1 mm. |
Calyces | colored as bracts, 15–25 mm; all 4 clefts subequal, 7–11 mm, 50–67% of calyx length; lobes linear, apices often slightly expanded, apex obtuse to acute. |
green to purple, margins green, white, or pink, 10–19 mm; abaxial, adaxial, and lateral clefts 3.5–7(–10) mm, 33–55% of calyx length, all 4 clefts subequal; lobes linear, lanceolate-elliptic, or narrowly triangular, apex acute to rounded. |
Filaments | glabrous. |
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Stigmas | black. |
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2n | = 20. |
= 24. |
Castilleja lineariloba |
Castilleja nana |
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Phenology | Flowering Apr–Jun(–Jul). | Flowering Jun–Aug. |
Habitat | Grasslands, moist meadows, swales, shores, forest openings. | Rocky or gravelly slopes, talus, ridges, fellfields, subalpine and alpine, often over granite. |
Elevation | 0–1600 m. (0–5200 ft.) | 1900–4300 m. (6200–14100 ft.) |
Distribution |
CA
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CA; NV; UT
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Discussion | Castilleja lineariloba is endemic to the western slope of the Sierra Nevada. Its chromosome number is 2n = 20, an apparent aneuploid reduction and documented by numerous counts. This diploid number is shared only with two very distantly related annual species endemic to central Mexico, C. gracilis Bentham and C. tenuifolia M. Martens & Galeotti. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja nana is limited to high elevations in the Sierra Nevada of California and the Great Basin ranges in central Nevada and western Utah. Plants with pink to purple inflorescences were described as C. lapidicola. Localized hybrid swarms between C. nana and C. viscidula are known from several mountain ranges in central and northern Nevada. Castilleja nana is sometimes confused with higher elevation forms of C. pilosa but can usually be distinguished from that species by the blackish patches on the sides of the corolla tube. Divergent populations in the central Sierra Nevada and adjacent White Mountains deserve further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 623. | FNA vol. 17, p. 633. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | Orthocarpus linearilobus, O. mariposanus | C. lapidicola |
Name authority | (Bentham) T. I. Chuang & Heckard: Syst. Bot. 16: 657. (1991) | Eastwood: Proc. Calif. Acad. Sci., ser. 3, 2: 289. (1902) — (as Castilleia) |
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