Castilleja levisecta |
Castilleja scabrida |
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golden Indian paintbrush, golden paintbrush |
rough Indian paintbrush, rough paintbrush |
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Habit | Herbs, perennial, 1–5 dm; from a woody caudex; with a taproot. | Herbs, perennial, 0.7–1.5(–2.2) dm; from a woody caudex; with a taproot or stout, branched roots. | ||||
Stems | few to many, erect, ± decumbent or creeping at base, unbranched, sometimes branched, hairs spreading, medium length and long, soft, mixed with short stipitate-glandular ones. |
several, decumbent, often sprawling, distally ascending, unbranched unless injured, rarely with small, leafy axillary shoots, hairs spreading, whitish, short, ± stiff, eglandular. |
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Leaves | green to purple or brown-tinged, linear-lanceolate proximally, oblong-ovate or -obovate distally, 0.8–5.2 cm, not fleshy, margins plane, distalmost sometimes ± wavy, involute, 3–7(–11)-lobed, apex obtuse; lobes erect to ascending, linear to linear-spatulate, very short, toothlike, usually arising from distal 1/3 of blade, apex rounded. |
gray-green, sometimes green or reddish purple, reduced and scalelike on proximal 10–25% of stem, linear, lanceolate, or narrowly elliptic, 1.5–4(–5) cm, not fleshy, margins plane, flat or involute, 0–3(–5)-lobed, apex acute; lobes spreading or spreading-ascending, linear or lanceolate, apex acute. |
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Inflorescences | 2.5–25 × 1–4 cm; bracts bright yellow throughout, or proximally greenish, distally bright yellow, sometimes deep yellow-orange, especially with age, oblong, elliptic, or obtuse to ovate, (0–)5–9(–13)-lobed, sometimes wavy-margined; lobes erect to ascending, oblong, short to medium length, arising above mid length, central lobe apex rounded, lateral ones rounded to acute. |
2.5–10 × 2.5–5 cm; bracts proximally green to deep greenish purple, distally bright red, sometimes brick red or orange-red, linear to lanceolate, 3–5(–7)-lobed; lobes spreading, linear-lanceolate to lanceolate, sometimes expanded near tip, long, arising near base and more distally, apex acute to obtuse. |
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Corollas | straight or slightly curved, 17–28 mm; tube 12–15 mm; beak exserted, adaxially green or greenish yellow, 6–8 mm; abaxial lip yellow or greenish, reduced, not inflated, 2–3 mm, 25–33(–50)% as long as beak; teeth ascending to erect, yellow, 0.5–1.5 mm. |
straight or slightly curved, 25–40(–45) mm; tube 11–24 mm; beak exserted, adaxially green to yellow, 10–17(–20) mm; abaxial lip green, reduced, visible through front cleft, 1.5–2.5 mm, 10–15% as long as beak; teeth incurved, green, 0.5–1 mm. |
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Calyces | distally yellow, 13–22 mm; abaxial and adaxial clefts 4–9.5 mm, 30–40% of calyx length, deeper than laterals, lateral 2.5–4.5 mm, ca. 25% of calyx length; lobes linear to narrowly oblong or narrowly lanceolate, apex obtuse, sometimes rounded to acute. |
colored as bracts, 18–33 mm; abaxial clefts 6–12 mm, adaxial 8–15 mm, clefts 40–60% of calyx length, deeper than laterals, lateral 2–6 mm, 15–25% of calyx length; lobes lanceolate or narrowly triangular, apex acute to acuminate. |
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2n | = 24. |
= 24. |
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Castilleja levisecta |
Castilleja scabrida |
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Phenology | Flowering Apr–Jul(–Sep). | |||||
Habitat | Dry meadows, upland prairie remnants, sunny headlands and coastal bluffs, rocky islands, often over glacial outwash or deposits. | |||||
Elevation | 0–400 m. (0–1300 ft.) | |||||
Distribution |
OR; WA; BC
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AZ; CO; NM; NV; UT
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Discussion | Castilleja levisecta is listed as threatened in the United States and endangered in Canada, where it is extremely rare. Most of its grassland habitat has been altered by development in the Puget Trough, and there are historical stations in the metro areas of what are now Victoria, Portland, and Seattle. For several decades, C. levisecta was considered extirpated from Oregon. However, recent reintroduction programs in Oregon and Washington have been very successful at reestablishing this species at several sites in the Willamette Valley. The bright yellow inflorescences often gradually age to a golden yellow color, unique in the genus. Castilleja levisecta is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Castilleja scabrida consists of two varieties distinguished by range and habitat differences, as well as by the morphological characters of the key. Both varieties are often confused with the similar and broadly sympatric C. chromosa but can be differentiated from the latter most easily by the presence of scalelike vestigial leaves on the proximal stems of C. scabrida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 621. | FNA vol. 17, p. 654. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Greenman: Bot. Gaz. 25: 268. (1898) — (as Castilleia) | Eastwood: Bull. Torrey Bot. Club 29: 523. (1902) — (as Castilleia) | ||||
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