Castilleja lemmonii |
Castilleja hyperborea |
|
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Lemmon's Indian paintbrush, Lemmon's paintbrush |
northern Indian paintbrush, northern paintbrush |
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Habit | Herbs, perennial, 0.8–2.5 dm; from a woody caudex; with slender, branching roots. | Herbs, perennial, 0.7–2.6 dm; from a many-headed, short, woody caudex; with a taproot. |
Stems | few to many, decumbent-based to erect, unbranched except for short, leafy axillary shoots, hairs sparse, spreading, medium length to long, soft and dense, short to medium length, stipitate-glandular. |
few, ascending, unbranched, sometimes branched, glabrate to sparsely hairy proximally, becoming densely hairy distally, proximal hairs appressed, white, short, soft, eglandular, distal hairs widely spreading to erect, white or yellowish, long to very long, soft, eglandular. |
Leaves | green or gray-green to purple (sometimes different on stems of same plant), linear-lanceolate, distal sometimes broadly lanceolate, 0.5–4 cm, not fleshy, margins plane, involute, 0(–3)-lobed, apex acute to acuminate; lobes ascending, linear to narrowly lanceolate, apex acute to acuminate, rarely obtuse. |
green to sometimes red-brown or dull purplish, linear to linear-lanceolate, 1.5–4 cm, not fleshy, margins plane, sometimes ± wavy, involute, (0–)3(–5)-lobed, apex acute to acuminate; lobes linear or linear-lanceolate, laterals ± divaricate, often abruptly up-curved from plane of main leaf blade, proximals usually 33–50% as long as leaf, apex acuminate to acute. |
Inflorescences | 2–12 × 1–3 cm; bracts greenish to dull purplish or brownish throughout, or proximally greenish to dull purplish, distally pink to purple or magenta, rarely white, ovate, broadly lanceolate, or oblong, (0–)3–5-lobed; lobes ascending to erect, lanceolate, medium length, arising above mid length, apex acute to rounded. |
2–11 × 1–2.5 cm; bracts greenish or pale yellow, rarely purplish or deep red, often with dull purplish veins on proximal bracts, or proximally greenish or pale yellow, distally whitish, yellow, or orange-yellow, proximal sometimes linear-lanceolate, distal broadly lanceolate to oblong or obovate, (0–)3–5(–7)-lobed; lobes ascending to erect, linear to lanceolate, short to medium length, arising above mid length, proximal bract apex acute, distal obtuse, rounded, or truncate. |
Corollas | slightly curved, 16–21 mm; tube 10–16 mm; abaxial lip sometimes partly exserted, beak usually exserted; beak adaxially green, 6–7 mm, margins red; abaxial lip greenish, inflated, pouches 3, shallow, central pouch shallowly grooved, visible through front cleft, 3–4 mm, 60% as long as beak; teeth erect, violet-purple or pink, 1–2.5 mm. |
slightly curved, 10–22 mm; tube 8–14 mm; teeth of abaxial lip sometimes slightly exserted, beak exserted or subequal; beak adaxially green to pale yellow, 5–8 mm; abaxial lip deep yellow or green, reduced, forming a small pouch visible through front cleft, 3–5 mm, 50–75% as long as beak; teeth ascending to erect, white or yellow, 1 mm. |
Calyces | proximally brown or dull magenta, sometimes green, distally colored as bracts, 12.5–18 mm; abaxial and adaxial clefts 5.5–10.5 mm, 40–65% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–15% of calyx length; lobes oblong, apex rounded. |
colored as bracts, proximally often paler, usually yellow throughout, 10–20 mm; abaxial and adaxial clefts 7–14 mm, 40–50% of calyx length, deeper than laterals, lateral 0.5–1.5 mm, 5–15% of calyx length; lobes triangular, apex acute to rounded. |
Stigmas | greenish to deep bluish purple. |
|
2n | = 24. |
= 24. |
Castilleja lemmonii |
Castilleja hyperborea |
|
Phenology | Flowering Jun–Aug. | Flowering May–Aug(–Sep). |
Habitat | Moist to wet meadows and flats, shorelines, open conifer forests, subalpine and alpine, often over granite. | Dry to mesic rocky tundra, slopes, ridges, barrens and meadows, openings in boreal forests, arctic to alpine. |
Elevation | 1500–3700 m. [4900–12100 ft.] | 0–2200 m. [0–7200 ft.] |
Distribution |
CA; NV
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AK; BC; NT; YT; e Asia (Russian Far East)
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Discussion | As delimited here, Castilleja lemmonii is endemic to the highlands of the Sierra Nevada in California and in adjacent Washoe County, Nevada. It differs from C. lassenensis, a plant of volcanic highlands around Mt. Lassen, which has consistently white corollas. Corollas are usually pink to purplish in C. lemmonii. Castilleja lemmonii also tends to have somewhat shorter lateral calyx clefts, though the two species overlap slightly in this character. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja hyperborea is widespread across boreal, alpine, and arctic habitats in western Canada and Alaska, extending into the Russian Far East. A rare form with light purple bracts was named as C. kuschei. Another form with particularly long hairs found in the Kluane Lake region of the southern Yukon was described as C. villosissima and is usually found on calcareous substrates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 620. | FNA vol. 17, p. 614. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. culbertsonii | C. kuschei, C. villosissima |
Name authority | A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 297. (1878) — (as Castilleia lemmoni) | Pennell: Proc. Acad. Nat. Sci. Philadelphia 86: 532. (1934) |
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