Castilleja lemmonii |
Castilleja dissitiflora |
|
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Lemmon's Indian paintbrush, Lemmon's paintbrush |
Mount Hamilton paintbrush, Mt. Hamilton Indian paintbrush |
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Habit | Herbs, perennial, 0.8–2.5 dm; from a woody caudex; with slender, branching roots. | Herbs, perennial, 1.8–4(–5) dm; from a woody caudex; with a taproot. |
Stems | few to many, decumbent-based to erect, unbranched except for short, leafy axillary shoots, hairs sparse, spreading, medium length to long, soft and dense, short to medium length, stipitate-glandular. |
several to many, erect to ascending, unbranched, sometimes branched, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones. |
Leaves | green or gray-green to purple (sometimes different on stems of same plant), linear-lanceolate, distal sometimes broadly lanceolate, 0.5–4 cm, not fleshy, margins plane, involute, 0(–3)-lobed, apex acute to acuminate; lobes ascending, linear to narrowly lanceolate, apex acute to acuminate, rarely obtuse. |
green, linear to narrowly or broadly lanceolate, (1–)3–5(–6) cm, not fleshy, margins wavy (obscure on many pressed specimens), involute, usually 0–3(–5)-lobed, apex broadly acute to rounded; lobes widely spreading, linear to narrowly lanceolate, apex acute to acuminate. |
Inflorescences | 2–12 × 1–3 cm; bracts greenish to dull purplish or brownish throughout, or proximally greenish to dull purplish, distally pink to purple or magenta, rarely white, ovate, broadly lanceolate, or oblong, (0–)3–5-lobed; lobes ascending to erect, lanceolate, medium length, arising above mid length, apex acute to rounded. |
2.5–10 × 2–5.5 cm; bracts proximally greenish, distally red to red-orange, narrowly lanceolate to broadly lanceolate, 3–5-lobed; lobes spreading, distal, if present, ascending, linear or narrowly oblanceolate to triangular, proximals long, arising below mid blade, distals short, sometimes mere teeth, near apex of central lobe, sometimes wavy-margined, apex obtuse to rounded, sometimes acute. |
Corollas | slightly curved, 16–21 mm; tube 10–16 mm; abaxial lip sometimes partly exserted, beak usually exserted; beak adaxially green, 6–7 mm, margins red; abaxial lip greenish, inflated, pouches 3, shallow, central pouch shallowly grooved, visible through front cleft, 3–4 mm, 60% as long as beak; teeth erect, violet-purple or pink, 1–2.5 mm. |
straight to slightly curved, 24–38 mm; tube 14–21 mm; beak exserted from calyx, adaxially green, 11–16.5(–18) mm; abaxial lip green, reduced, visible or not through deep front cleft in calyx, 2 mm, 13% as long as beak; teeth incurved, green, 1 mm. |
Calyces | proximally brown or dull magenta, sometimes green, distally colored as bracts, 12.5–18 mm; abaxial and adaxial clefts 5.5–10.5 mm, 40–65% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–15% of calyx length; lobes oblong, apex rounded. |
whitish with green veins or green, sometimes purple, distally same color as bracts, sometimes with yellowish band below colored apices, 20–26(–29) mm; abaxial clefts (8–)13–16(–19) mm, adaxial 7–12(–14) mm, clefts 35–50% of calyx length, deeper than laterals, lateral 2–6(–8) mm, 10–30% of calyx length; lobes linear-lanceolate to lanceolate, apex obtuse to acute. |
Stigmas | greenish to deep bluish purple. |
|
2n | = 24. |
= 48. |
Castilleja lemmonii |
Castilleja dissitiflora |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Moist to wet meadows and flats, shorelines, open conifer forests, subalpine and alpine, often over granite. | Sagebrush slopes often rocky, montane to subalpine. |
Elevation | 1500–3700 m. [4900–12100 ft.] | 1900–3300 m. [6200–10800 ft.] |
Distribution |
CA; NV
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NV
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Discussion | As delimited here, Castilleja lemmonii is endemic to the highlands of the Sierra Nevada in California and in adjacent Washoe County, Nevada. It differs from C. lassenensis, a plant of volcanic highlands around Mt. Lassen, which has consistently white corollas. Corollas are usually pink to purplish in C. lemmonii. Castilleja lemmonii also tends to have somewhat shorter lateral calyx clefts, though the two species overlap slightly in this character. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja dissitiflora is endemic to several mountain ranges in central and eastern Nevada, in the upper montane and lower subalpine zones. It has the deep abaxial calyx cleft of C. linariifolia and the stipitate-glandular, wavy-margined leaves of C. applegatei var. pinetorum. Based on morphological data, Holmgren suggested that it is an allopolyploid derived from hybridization of C. applegatei var. pinetorum and C. linariifolia. His proposal is plausible and should be further tested. Castilleja dissitiflora is a tetraploid, while both putative parental species have at least some diploid populations. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 620. | FNA vol. 17, p. 603. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. culbertsonii | |
Name authority | A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 297. (1878) — (as Castilleia lemmoni) | N. H. Holmgren: Mem. New York Bot. Gard. 21(4): 46, figs. 6–8. (1971) |
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