Castilleja lemmonii |
Castilleja chrysantha |
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Lemmon's Indian paintbrush, Lemmon's paintbrush |
Wallowa paintbrush, yellow paintbrush, yellow Wallowa Indian paintbrush, yellowish or common Wallowa paintbrush |
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Habit | Herbs, perennial, 0.8–2.5 dm; from a woody caudex; with slender, branching roots. | Herbs, perennial, (0.5–)1–2(–5) dm; from a woody caudex; with a taproot. |
Stems | few to many, decumbent-based to erect, unbranched except for short, leafy axillary shoots, hairs sparse, spreading, medium length to long, soft and dense, short to medium length, stipitate-glandular. |
few to many, erect or ascending, often decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, mixed with dense, shorter stipitate-glandular ones. |
Leaves | green or gray-green to purple (sometimes different on stems of same plant), linear-lanceolate, distal sometimes broadly lanceolate, 0.5–4 cm, not fleshy, margins plane, involute, 0(–3)-lobed, apex acute to acuminate; lobes ascending, linear to narrowly lanceolate, apex acute to acuminate, rarely obtuse. |
green or purple-tinged, sometimes deep purple, broadly lanceolate, sometimes linear-lanceolate or narrowly oblong, 1.5–4.8 cm, not fleshy, margins plane, flat or involute, 0–3-lobed, apex acute; lobes ascending or spreading, linear to narrowly linear or narrowly lanceolate, short, apex acuminate. |
Inflorescences | 2–12 × 1–3 cm; bracts greenish to dull purplish or brownish throughout, or proximally greenish to dull purplish, distally pink to purple or magenta, rarely white, ovate, broadly lanceolate, or oblong, (0–)3–5-lobed; lobes ascending to erect, lanceolate, medium length, arising above mid length, apex acute to rounded. |
3–17 × 1–3 cm; bracts greenish or pale yellow-green throughout, or proximally greenish or pale yellow-green, distally pale yellow to whitish, sometimes pink-purple, or pale, dull purplish, sometimes aging pink or yellow, often infused with light purple, rarely pink, ovate to broadly acute, (0–)3-lobed; lobes ascending, linear-lanceolate, oblong, or triangular, short, arising below mid length, apex acute to obtuse. |
Corollas | slightly curved, 16–21 mm; tube 10–16 mm; abaxial lip sometimes partly exserted, beak usually exserted; beak adaxially green, 6–7 mm, margins red; abaxial lip greenish, inflated, pouches 3, shallow, central pouch shallowly grooved, visible through front cleft, 3–4 mm, 60% as long as beak; teeth erect, violet-purple or pink, 1–2.5 mm. |
straight, (16–)20–25 mm; tube 15–18 mm; beak slightly exserted, adaxially green to yellow, 5.5–8.5 mm; abaxial lip white, green and white, pink, or purple, ± prominent, appressed (proximally scarcely or not pouched), 3–5 mm, 67% as long as beak; teeth erect, white or pink with some purple or red, 1.5–2.5 mm. |
Calyces | proximally brown or dull magenta, sometimes green, distally colored as bracts, 12.5–18 mm; abaxial and adaxial clefts 5.5–10.5 mm, 40–65% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–15% of calyx length; lobes oblong, apex rounded. |
proximally green, pale with green veins, purple-tinged green, or purple, distally pale yellow, white, or purplish, 12–20 mm; abaxial and adaxial clefts 8–12 mm, 50% of calyx length, deeper than laterals, lateral 0.5–3 mm, 5–15% of calyx length; lobes short-triangular, apex obtuse, rounded, or truncate. |
Stigmas | greenish to deep bluish purple. |
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2n | = 24. |
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Castilleja lemmonii |
Castilleja chrysantha |
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Phenology | Flowering Jun–Aug. | Flowering May–Oct. |
Habitat | Moist to wet meadows and flats, shorelines, open conifer forests, subalpine and alpine, often over granite. | Flat, mesic meadows, dry talus and ridges, montane to alpine. |
Elevation | 1500–3700 m. [4900–12100 ft.] | 1100–2800 m. [3600–9200 ft.] |
Distribution |
CA; NV
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OR
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Discussion | As delimited here, Castilleja lemmonii is endemic to the highlands of the Sierra Nevada in California and in adjacent Washoe County, Nevada. It differs from C. lassenensis, a plant of volcanic highlands around Mt. Lassen, which has consistently white corollas. Corollas are usually pink to purplish in C. lemmonii. Castilleja lemmonii also tends to have somewhat shorter lateral calyx clefts, though the two species overlap slightly in this character. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja chrysantha is endemic to the mountains of northeastern Oregon, and its patterns of variation need further study. Most plants in the Blue Mountains are taller, more erect, and tolerate lower elevation habitats than populations around the type locality in the Wallowa Mountains. Plants with purplish inflorescences and longer hairs were described as C. ownbeyana and tend to favor drier talus and ridges than plants like the type, found in flat, mesic, montane to subalpine meadows. Hybrids between C. chrysantha and C. fraterna occur in Wallowa County. In the Wallowa Mountains, a recurrent and variable hybrid form between C. chrysantha and C. rhexiifolia was described as C. wallowensis Pennell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 620. | FNA vol. 17, p. 596. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. culbertsonii | C. ownbeyana |
Name authority | A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 297. (1878) — (as Castilleia lemmoni) | Greenman: Bot. Gaz. 48: 146. (1909) |
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