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Sierra woolly Indian paintbrush, Sierra woolly paintbrush, woolly paintbrush

wavyleaf Indian paintbrush

Habit Subshrubs or herbs, perennial, 1.8–9(–10) dm; from a woody caudex; with a thick, woody taproot or system to thick, branched roots. Herbs, perennial, 1.4–8 dm; from a woody caudex; with a taproot.
Stems

few to many, erect to ascending, unbranched, sometimes branched distally, sometimes with leafy axillary shoots, lanate to woolly, hairs dense, spreading to weakly appressed, white to yellowish, medium length, soft, moderately branched or unbranched in different populations, sometimes mixed, eglandular, rarely short stipitate-glandular, mostly obscuring surface.

many, erect, unbranched or branched, sometimes with small, leafy axillary branches, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones.

Leaves

green to purple, often hidden by matted, white hairs, linear to narrowly oblong or narrowly lanceolate, (1–)2–7(–10) cm, not fleshy, margins plane, flat, sometimes involute, 0–3-lobed, apex rounded, sometimes acute;

lobes spreading, linear or narrowly oblong, apex rounded to acute.

green or purple-tinged, lanceolate to linear-oblong, 0.8–6.1 cm, not fleshy, margins wavy, involute, 0(–3)-lobed, apex rounded to acute;

lobes spreading-ascending, narrowly lanceolate, apex acute to acuminate.

Inflorescences

2–19 × 1.5–5 cm;

bracts proximally pale greenish to greenish gray, distally bright red to orange-red, sometimes pinkish, magenta, or salmon, rarely yellow, oblong to narrowly lanceolate or narrowly oblanceolate, deeply 0–3(–5)-lobed;

lobes spreading, oblanceolate or linear, long, arising below mid length, distal bract apex usually rounded.

3–34 × 1.5–3.5 cm;

flowers usually distichous, remote, except distalmost;

bracts proximally greenish, distally greenish, reddish, orange-red, or dull orange, rarely yellow, proximal bracts frequently greenish throughout, lanceolate to narrowly ovate or narrowly oblong-elliptic, 0(–3)-lobed, proximal wavy-margined;

lobes ascending, triangular, short, arising near apex, apex acute to obtuse.

Corollas

curved in proximal 1/3, 23–35(–42) mm;

tube 12–17 mm;

beak exserted, adaxially greenish to yellowish, 11–22 mm;

abaxial lip dark green, reduced, 1–4 mm, ca. 20% as long as beak;

teeth incurved, reduced, green, 0.5–2.5 mm.

± curved, 25–38 mm;

tube 14–18 mm;

beak and often abaxial lip exserted;

beak adaxially dull orange or dull red, rarely yellow, 14–19 mm;

abaxial lip slightly curved, green, red, whitish, or yellow, not inflated, 2–2.5 mm, 15% as long as beak;

teeth incurved, deep green to reddish, 1–1.5 mm.

Calyces

proximally whitish to green, central band whitish, distal 1/3 colored as bract lobes, sometimes lighter or deeper, 15–29 mm;

abaxial and adaxial clefts 6–14 mm, ca. 50% of calyx length, deeper than laterals, lateral 0–5 mm, 0–5(–15)% of calyx length;

lobes oblong to broadly triangular, apex rounded, truncate, or emarginate.

proximally whitish, green, or purple, distally colored as bracts, 9–18 mm;

abaxial clefts 6–6.5 mm, adaxial 7–9 mm, clefts 33+% of calyx length, deeper than laterals, lateral 1.5–4 mm, ca. 10% of calyx length;

lobes linear-lanceolate, apex acute.

2n

= 24.

= 24.

Castilleja lanata

Castilleja disticha

Phenology Flowering year-round. Flowering (May–)Jun–Aug(–Sep).
Habitat Dry rocky slopes, ridges, canyons, flats, valleys, montane. Open conifer forests, rocky or sandy slopes, montane to subalpine.
Elevation 300–2300 m. (1000–7500 ft.) 1600–3000 m. (5200–9800 ft.)
Distribution
from FNA
AZ; NM; TX; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Most records of Castilleja lanata are from the southern third of Arizona, New Mexico, and southwestern Texas. Plants sometimes referred to C. lanata from the Channel Islands of California are assigned here to C. hololeuca, an insular endemic. In northern Mexico, C. lanata extends from San Luis Potosí and Nuevo León westward to Sonora, with a disjunct distribution in central Baja California, where it overlaps the southern limits of a related species, C. foliolosa. Castilleja lanata inhabits a wide spectrum of soils and elevations and does not co-occur with the closely related C. galehintoniae in Nuevo León, where the latter is limited to limestone and gypsum substrates in a narrow elevational range. Hybrids of C. lanata with C. sessiliflora are known from Pecos County, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Castilleja disticha is limited to the central and southern Sierra Nevada. Although similar to C. applegatei and C. martini, C. disticha is distinctive and unique in its long, colorful, and highly exserted corollas with the beak exceeding the tube in length, calyces more brightly colored than the often unlobed bracts, and tall, strongly distichous growth form.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 618. FNA vol. 17, p. 603.
Parent taxa Orobanchaceae > Castilleja Orobanchaceae > Castilleja
Sibling taxa
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. disticha, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. martini, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. montigena, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurascens, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. wootonii, C. xanthotricha
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lanata, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. martini, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. montigena, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurascens, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. wootonii, C. xanthotricha
Synonyms C. applegatei subsp. disticha
Name authority A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 118. (1859) Eastwood: Proc. Calif. Acad. Sci., ser. 3, 2: 289. (1902) — (as Castilleia)
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