Castilleja lanata |
Castilleja applegatei |
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Sierra woolly Indian paintbrush, Sierra woolly paintbrush, woolly paintbrush |
Applegate's Indian paintbrush, Applegate's paintbrush, wavy leaf paintbrush, wavy-leaf Indian paintbrush, waxy-leaf Indian paintbrush |
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Habit | Subshrubs or herbs, perennial, 1.8–9(–10) dm; from a woody caudex; with a thick, woody taproot or system to thick, branched roots. | Herbs, perennial, (0.8–)1–5(–6) dm; from a branched, woody caudex; with a taproot. | ||||||||||||
Stems | few to many, erect to ascending, unbranched, sometimes branched distally, sometimes with leafy axillary shoots, lanate to woolly, hairs dense, spreading to weakly appressed, white to yellowish, medium length, soft, moderately branched or unbranched in different populations, sometimes mixed, eglandular, rarely short stipitate-glandular, mostly obscuring surface. |
few to many, erect, ascending, or decumbent, unbranched or branched, hairs sparse to dense, spreading, long, soft to ± stiff, eglandular, mixed with shorter stipitate-glandular ones. |
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Leaves | green to purple, often hidden by matted, white hairs, linear to narrowly oblong or narrowly lanceolate, (1–)2–7(–10) cm, not fleshy, margins plane, flat, sometimes involute, 0–3-lobed, apex rounded, sometimes acute; lobes spreading, linear or narrowly oblong, apex rounded to acute. |
green to purplish or brown, linear to broadly lanceolate, sometimes ovate, 1–6 cm, not fleshy, margins wavy, involute, (0–)3(–5)-lobed, apex rounded or narrowly acute to acuminate; lobes spreading or ascending, lanceolate to broadly lanceolate, often short, apex acute to rounded. |
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Inflorescences | 2–19 × 1.5–5 cm; bracts proximally pale greenish to greenish gray, distally bright red to orange-red, sometimes pinkish, magenta, or salmon, rarely yellow, oblong to narrowly lanceolate or narrowly oblanceolate, deeply 0–3(–5)-lobed; lobes spreading, oblanceolate or linear, long, arising below mid length, distal bract apex usually rounded. |
2–12(–21 in fruit) × 1–5 cm; bracts proximally green to dull purplish brown, distally red, red-orange, or scarlet, sometimes orange, white, or yellow, rarely with a narrow yellowish band medially, lanceolate, broadly lanceolate, oblong, or lanceolate-ovate, 3–5(–7)-lobed; lobes spreading to ascending, linear, sometimes expanded near tip, long, arising from ca. mid length, central lobe apex obtuse to rounded, sometimes expanded, others acute, rounded, obtuse, or acuminate. |
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Corollas | curved in proximal 1/3, 23–35(–42) mm; tube 12–17 mm; beak exserted, adaxially greenish to yellowish, 11–22 mm; abaxial lip dark green, reduced, 1–4 mm, ca. 20% as long as beak; teeth incurved, reduced, green, 0.5–2.5 mm. |
straight or curved in proximal 1/3, 16–35(–41) mm; tube 9–22 mm; beak usually long-exserted, adaxially green, yellow-green, or yellow, 6–20 mm; abaxial lip deep green to whitish or yellow, reduced, inconspicuous, protuberant, thickened, included or exserted, 1–3 mm, 5–20% as long as beak; teeth ascending or incurved, deep green to yellow, 0.5–1 mm. |
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Calyces | proximally whitish to green, central band whitish, distal 1/3 colored as bract lobes, sometimes lighter or deeper, 15–29 mm; abaxial and adaxial clefts 6–14 mm, ca. 50% of calyx length, deeper than laterals, lateral 0–5 mm, 0–5(–15)% of calyx length; lobes oblong to broadly triangular, apex rounded, truncate, or emarginate. |
proximally green or whitish, sometimes yellow, lobes colored as bract lobes, sometimes with a yellow band between proximal and distal portions, 13–25 mm; abaxial and adaxial clefts 4–14 mm, 33–50% of calyx length, deeper than laterals, lateral 2.5–8 mm, 12–50% of calyx length; lobes lanceolate-acuminate, narrowly oblong, or narrowly triangular, apex acuminate, acute, or obtuse. |
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2n | = 24. |
= 24, 48. |
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Castilleja lanata |
Castilleja applegatei |
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Phenology | Flowering year-round. | |||||||||||||
Habitat | Dry rocky slopes, ridges, canyons, flats, valleys, montane. | |||||||||||||
Elevation | 300–2300 m. (1000–7500 ft.) | |||||||||||||
Distribution |
AZ; NM; TX; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
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CA; ID; NV; OR; UT; WY
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Discussion | Most records of Castilleja lanata are from the southern third of Arizona, New Mexico, and southwestern Texas. Plants sometimes referred to C. lanata from the Channel Islands of California are assigned here to C. hololeuca, an insular endemic. In northern Mexico, C. lanata extends from San Luis Potosí and Nuevo León westward to Sonora, with a disjunct distribution in central Baja California, where it overlaps the southern limits of a related species, C. foliolosa. Castilleja lanata inhabits a wide spectrum of soils and elevations and does not co-occur with the closely related C. galehintoniae in Nuevo León, where the latter is limited to limestone and gypsum substrates in a narrow elevational range. Hybrids of C. lanata with C. sessiliflora are known from Pecos County, Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Castilleja applegatei is a widespread and often common species, with complex patterns of variation and several common but inconstant color forms, especially in var. pinetorum. Castilleja disticha and C. martini, although sometimes included as subspecies within C. applegatei, are treated as species here. Both are more morphologically divergent and more easily distinguished from typical C. applegatei than are the four varieties accepted here. Variety pinetorum is the most widespread form and also occurs over a wider range of elevations than the other three varieties, which are primarily montane to subalpine and occasionally alpine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 618. | FNA vol. 17, p. 587. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Name authority | A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 118. (1859) | Fernald: Erythea 6: 49. (1898) — (as Castilleia) | ||||||||||||
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