Castilleja lacera |
Castilleja pilosa |
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cut-leaf owl's clover, cut-leaf paintbrush, cutleaf Indian paintbrush, foothill owl's clover |
hairy paintbrush, parrot-head Indian paintbrush, pilose paintbrush |
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Habit | Herbs, annual, 0.5–4 dm; with fibrous roots. | Herbs, perennial, (0.7–)1.2–3.5(–4.4) dm; from a woody caudex; with a stout taproot. | ||||||||
Stems | solitary, erect, unbranched or branched, hairs spreading, long, soft, scattered among more numerous, medium length, stipitate-glandular ones. |
several to many, ascending to erect, sometimes short-decumbent, branched or unbranched, sometimes with short, leafy axillary shoots, hairs moderately dense, retrorse or curved to spreading, straight, curly, or ± wavy, medium length to long, soft to stiff, eglandular. |
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Leaves | green or purplish, linear to narrowly lanceolate, 1–5 cm, not fleshy, margins plane, flat, 0–5(–7)-lobed, apex acuminate; lobes spreading to ascending, linear, apex acuminate to acute. |
green to purple, linear to lanceolate, 1–5.5(–8) cm, not fleshy, margins plane to ± wavy, involute, 0–5(–7)-lobed, apex acuminate to obtuse; lobes widely spreading to ascending-spreading, linear to filiform, apex acute or obtuse. |
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Inflorescences | (1.5–)3–14 × 2–3 cm; bracts green throughout, sometimes proximally green, distally white on apices, lanceolate to ovate, 3–7-lobed; lobes spreading to ascending, linear to narrowly lanceolate, long, arising below mid length, apex obtuse to acute. |
(2–)3.5–16 × 1–3.5 cm; bracts light green, green, yellow-green, light purple, purple, light dusky pink, salmon, or reddish brown throughout, or these colors proximally, distally or on distal margins white, pale yellow, yellow, pale salmon, or buff, sometimes becoming reddish purple with age, lanceolate to broadly lanceolate or narrowly ovate, 3–5(–9)-lobed, often wavy-margined; lobes spreading to erect or ascending, linear to oblanceolate, short to long, arising near or above mid length, sometimes wavy-margined, central lobe apex obtuse to rounded or truncate, sometimes acute, lateral ones acute to obtuse. |
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Corollas | straight, 10–22 mm; tube 8–15 mm; abaxial lip and beak exserted; beak adaxially yellow to greenish, 3–6 mm, densely puberulent; abaxial lip yellow with purple dots at base, inflated, pouches 3, central pouch slightly 2-lobed, pouches 4–8 mm wide, 3–6 mm deep, side pouches curving up a little at tip, 2–5 mm, 75–95% as long as beak; teeth erect, white or yellow, 0.5–2 mm. |
straight, 14–23 mm; tube 10–15; abaxial lip sometimes partially exserted, beak exserted; beak adaxially green or yellow-green, 3–7 mm; abaxial lip proximally green, pale yellow, pale or bright pink, or deep purple, distally white to pink or purplish, inflated, pouches 3, deeply furrowed, 2.5–8 mm, 50–100% as long as beak; teeth erect, green, white, buff, pink, or pale yellow, 1–2 mm. |
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Calyces | light green, lobes green, 7–13 mm; abaxial and adaxial clefts 3.5–8 mm, 50–67% of calyx length, lateral 2.5–5 mm, ca. 40% of calyx length; lobes narrowly to broadly lanceolate, apex acute to acuminate. |
proximally whitish to pale green, distally green, whitish, pink, or yellowish, 9–28 mm; all 4 clefts subequal, 2–12 mm, 45–55% of calyx length; lobes linear to narrowly lanceolate or narrowly triangular, rarely deltoid, apex acute, rarely rounded. |
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Stigmas | equal to or slightly exserted from beak. |
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2n | = 22, 24. |
= 24, 48, 96. |
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Castilleja lacera |
Castilleja pilosa |
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Phenology | Flowering (Mar–)Apr–Jul(–Aug). | |||||||||
Habitat | Grasslands, meadows, moist flats, vernal pool margins, moist forest openings, serpentine slopes and ledges, roadsides. | |||||||||
Elevation | 0–2700 m. (0–8900 ft.) | |||||||||
Distribution |
CA; OR
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CA; ID; MT; NV; OR; WY
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Discussion | Castilleja lacera is found in a wide range of elevations in the central and northern Sierra Nevada region and in the Siskiyou Mountains region of northwestern California and southwestern Oregon. Reports from the Coast Ranges north of the San Francisco Bay region and south of the Siskiyou region in western California are referable to other yellow-flowered annuals, including C. ambigua, C. rubicundula var. lithospermoides, Triphysaria eriantha subsp. eriantha, and T. versicolor subsp. faucibarbata. Although most similar to C. rubicundula, C. lacera is somewhat smaller in stature and flower size. It is also easily confused with yellow-flowered populations of C. tenuis, which has smaller flowers and an included stigma. Two chromosome numbers are known for this species, the more northern populations being diploid, and those to the south having an apparently aneuploid count of 2n = 22, which is unique in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Castilleja pilosa is a widespread and variable complex similar in growth form and coloration to C. pallescens and related species. However, C. pilosa is distinguished with relative ease by its subequally divided calyces. Plants of the C. pallescens complex have very shallow lateral calyx lobes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 617. | FNA vol. 17, p. 644. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Orthocarpus lacerus | Orthocarpus pilosus | ||||||||
Name authority | (Bentham) T. I. Chuang & Heckard: Syst. Bot. 16: 657. (1991) | (S. Watson) Rydberg: Mem. New York Bot. Gard. 1: 361. (1900) | ||||||||
Web links |