Castilleja hispida |
Castilleja lanata |
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harsh Indian paintbrush, harsh paintbrush, hispid or harsh or bristly paintbrush |
Sierra woolly Indian paintbrush, Sierra woolly paintbrush, woolly paintbrush |
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Habit | Herbs, perennial, 1.3–5(–6) dm; from a woody caudex; with a taproot. | Subshrubs or herbs, perennial, 1.8–9(–10) dm; from a woody caudex; with a thick, woody taproot or system to thick, branched roots. | ||||
Stems | few to many, erect or ascending, unbranched, sometimes with inconspicuous, short, leafy axillary shoots, hairs spreading to erect, long, soft to stiff, mixed with shorter stipitate-glandular ones. |
few to many, erect to ascending, unbranched, sometimes branched distally, sometimes with leafy axillary shoots, lanate to woolly, hairs dense, spreading to weakly appressed, white to yellowish, medium length, soft, moderately branched or unbranched in different populations, sometimes mixed, eglandular, rarely short stipitate-glandular, mostly obscuring surface. |
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Leaves | green, sometimes purple-tinged, margins sometimes red-brown, linear or narrowly to broadly lanceolate to narrowly oblong, oblanceolate, or ovate, 1–8.5 cm, not fleshy, margins plane or wavy, involute or flat, (0–)3–5(–7)-lobed, apex acute to rounded or acuminate; lobes ascending, linear to narrowly lanceolate, much narrower than mid blade, apex acute to rounded. |
green to purple, often hidden by matted, white hairs, linear to narrowly oblong or narrowly lanceolate, (1–)2–7(–10) cm, not fleshy, margins plane, flat, sometimes involute, 0–3-lobed, apex rounded, sometimes acute; lobes spreading, linear or narrowly oblong, apex rounded to acute. |
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Inflorescences | 3–16(–30 in fruit) × 2–5 cm; bracts proximally greenish to dull reddish purple, distally red to orange or yellow, sometimes crimson, scarlet, orange-red, red-orange, or burnt orange, often becoming paler and/or duller with age, lanceolate to oblong, ovate, or obovate, 3–5(–11)-lobed; lobes spreading to ascending, linear to oblong or narrowly lanceolate, medium length or long, arising at or above mid length, central lobe apex obtuse to rounded or truncate, sometimes emarginate, truncate, or acute, lateral ones acute to obtuse. |
2–19 × 1.5–5 cm; bracts proximally pale greenish to greenish gray, distally bright red to orange-red, sometimes pinkish, magenta, or salmon, rarely yellow, oblong to narrowly lanceolate or narrowly oblanceolate, deeply 0–3(–5)-lobed; lobes spreading, oblanceolate or linear, long, arising below mid length, distal bract apex usually rounded. |
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Corollas | slightly curved, 17–38 mm; tube 12–18 mm; beak exserted, adaxially green, sometimes yellowish, rarely red-brown, 9–20 mm; abaxial lip ascending, deep green, reduced, curved, 0.5–3 mm, to 10–33% length of beak; teeth incurved, reduced, green, 0.5–1.2 mm. |
curved in proximal 1/3, 23–35(–42) mm; tube 12–17 mm; beak exserted, adaxially greenish to yellowish, 11–22 mm; abaxial lip dark green, reduced, 1–4 mm, ca. 20% as long as beak; teeth incurved, reduced, green, 0.5–2.5 mm. |
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Calyces | colored as bracts, sometimes with a yellow band proximal to red to orange apices, or ca. 1/2 yellowish and 1/2 reddish, 12–35 mm; abaxial and adaxial clefts 7–12 mm, 33–65% of calyx length, deeper than laterals, lateral 2–7 mm, 15–30% of calyx length; lobes triangular, linear, or oblong to lanceolate, apex acute or obtuse to rounded. |
proximally whitish to green, central band whitish, distal 1/3 colored as bract lobes, sometimes lighter or deeper, 15–29 mm; abaxial and adaxial clefts 6–14 mm, ca. 50% of calyx length, deeper than laterals, lateral 0–5 mm, 0–5(–15)% of calyx length; lobes oblong to broadly triangular, apex rounded, truncate, or emarginate. |
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2n | = 24, 48, 96. |
= 24. |
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Castilleja hispida |
Castilleja lanata |
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Phenology | Flowering year-round. | |||||
Habitat | Dry rocky slopes, ridges, canyons, flats, valleys, montane. | |||||
Elevation | 300–2300 m. (1000–7500 ft.) | |||||
Distribution |
ID; MT; OR; WA; AB; BC
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AZ; NM; TX; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
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Discussion | Varieties 2 (2 in the flora). Castilleja hispida is likely related to C. chromosa, which replaces it geographically to the southeast. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most records of Castilleja lanata are from the southern third of Arizona, New Mexico, and southwestern Texas. Plants sometimes referred to C. lanata from the Channel Islands of California are assigned here to C. hololeuca, an insular endemic. In northern Mexico, C. lanata extends from San Luis Potosí and Nuevo León westward to Sonora, with a disjunct distribution in central Baja California, where it overlaps the southern limits of a related species, C. foliolosa. Castilleja lanata inhabits a wide spectrum of soils and elevations and does not co-occur with the closely related C. galehintoniae in Nuevo León, where the latter is limited to limestone and gypsum substrates in a narrow elevational range. Hybrids of C. lanata with C. sessiliflora are known from Pecos County, Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 612. | FNA vol. 17, p. 618. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 105. (1838) | A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 118. (1859) | ||||
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