Castilleja hispida |
Castilleja exserta |
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harsh Indian paintbrush, harsh paintbrush, hispid or harsh or bristly paintbrush |
castilleja exserta, escobita, exserted Indian paintbrush, owl's clover, purple owl's-clover, purple painted-cup |
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Habit | Herbs, perennial, 1.3–5(–6) dm; from a woody caudex; with a taproot. | Herbs, annual, 0.1–4.5 dm; with fibrous roots. | ||||||||||||
Stems | few to many, erect or ascending, unbranched, sometimes with inconspicuous, short, leafy axillary shoots, hairs spreading to erect, long, soft to stiff, mixed with shorter stipitate-glandular ones. |
solitary, erect to ascending, unbranched or diffusely branched from near base, hairs spreading, medium length and long, stiff, mixed with short stipitate-glandular ones. |
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Leaves | green, sometimes purple-tinged, margins sometimes red-brown, linear or narrowly to broadly lanceolate to narrowly oblong, oblanceolate, or ovate, 1–8.5 cm, not fleshy, margins plane or wavy, involute or flat, (0–)3–5(–7)-lobed, apex acute to rounded or acuminate; lobes ascending, linear to narrowly lanceolate, much narrower than mid blade, apex acute to rounded. |
green, sometimes purple to brownish, linear or ovate to orbicular in outline, (0.8–)1–5(–7.7) cm, not fleshy, margins plane, involute, (0–)3–9(–11)-lobed, sometimes with secondary lobing, apex acuminate to rounded or acute; lobes spreading, filiform or linear to narrowly spatulate, apex acute to acuminate or rounded. |
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Inflorescences | 3–16(–30 in fruit) × 2–5 cm; bracts proximally greenish to dull reddish purple, distally red to orange or yellow, sometimes crimson, scarlet, orange-red, red-orange, or burnt orange, often becoming paler and/or duller with age, lanceolate to oblong, ovate, or obovate, 3–5(–11)-lobed; lobes spreading to ascending, linear to oblong or narrowly lanceolate, medium length or long, arising at or above mid length, central lobe apex obtuse to rounded or truncate, sometimes emarginate, truncate, or acute, lateral ones acute to obtuse. |
1.5–20 × 2–4 cm; bracts proximally greenish, dark purple, brownish purple, or white, distally pink, lavender, magenta, light purple, or white on lobe apices, lanceolate to elliptic or narrowly ovate, (3–)5(–9)-lobed, often with 2–4 secondary lobes; lobes ascending to spreading, linear to filiform or narrowly spatulate, medium length to long, arising below or above mid length, apex rounded to acute. |
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Corollas | slightly curved, 17–38 mm; tube 12–18 mm; beak exserted, adaxially green, sometimes yellowish, rarely red-brown, 9–20 mm; abaxial lip ascending, deep green, reduced, curved, 0.5–3 mm, to 10–33% length of beak; teeth incurved, reduced, green, 0.5–1.2 mm. |
straight, 12–33 mm; tube 7–20 mm; beak slightly exserted, hooked near apex, adaxially pink-purple to magenta, rarely white, 5–13 mm, margins colored as bracts, densely villous-hairy; abaxial lip proximally pink, purple, or magenta, rarely yellow or white, with maroon or deep purple distal to that and white to yellow or pink distally, often with purple spots, distal pale color often aging deep pink or deep red, strongly inflated, pouches 3, 3–8 mm wide, 3–4 mm deep, 3–9 mm, 67–80% as long as beak; teeth erect, white, yellow, or purple, often with purple or maroon spots, 0.5–2 mm. |
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Calyces | colored as bracts, sometimes with a yellow band proximal to red to orange apices, or ca. 1/2 yellowish and 1/2 reddish, 12–35 mm; abaxial and adaxial clefts 7–12 mm, 33–65% of calyx length, deeper than laterals, lateral 2–7 mm, 15–30% of calyx length; lobes triangular, linear, or oblong to lanceolate, apex acute or obtuse to rounded. |
colored as bracts, 10–26 mm; abaxial clefts 4–12 mm, adaxial 9–18 mm, abaxial ca. 50% of calyx length, adaxial ca. 67% of calyx length, at least adaxial deeper than others, lateral 2.5–9 mm, 15–45% of calyx length; lobes linear (to narrowly oblanceolate), apex rounded to acute. |
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Filaments | with spreading, long, soft hairs. |
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2n | = 24, 48, 96. |
= 24. |
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Castilleja hispida |
Castilleja exserta |
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Distribution |
ID; MT; OR; WA; AB; BC
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AZ; CA; NM; nw Mexico
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Discussion | Varieties 2 (2 in the flora). Castilleja hispida is likely related to C. chromosa, which replaces it geographically to the southeast. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). In addition to the characters in the key, Castilleja exserta is distinguished from the similar C. densiflora by its conspicuously hairy and apically hooked beak. As a result, the capitate stigma is exserted more or less horizontally from the corolla beak. In contrast, C. densiflora has an unhooked, inconspicuously puberulent beak, from which the stigma emerges more vertically. Castilleja exserta hybridizes with C. attenuata in southern California and with C. lineariloba in central California, and it reportedly crosses occasionally with C. densiflora in southern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 612. | FNA vol. 17, p. 606. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Synonyms | Orthocarpus exsertus | |||||||||||||
Name authority | Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 105. (1838) | (A. Heller) T. I. Chuang & Heckard: Syst. Bot. 16: 657. (1991) | ||||||||||||
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