Castilleja hispida |
Castilleja elmeri |
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harsh Indian paintbrush, harsh paintbrush, hispid or harsh or bristly paintbrush |
Elmer's Indian paintbrush, Elmer's paintbrush, Wenatchee Indian paintbrush, Wenatchee paintbrush |
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Habit | Herbs, perennial, 1.3–5(–6) dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.1–4.2 dm; from a many-headed, woody caudex; with a stout taproot. | ||||
Stems | few to many, erect or ascending, unbranched, sometimes with inconspicuous, short, leafy axillary shoots, hairs spreading to erect, long, soft to stiff, mixed with shorter stipitate-glandular ones. |
solitary or few to many, erect to ascending, sometimes slightly curved at base, unbranched, rarely branched, hairs moderately dense, spreading, medium length, soft, mixed with shorter eglandular and stipitate-glandular ones, at least on distal 1/2 of stem. |
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Leaves | green, sometimes purple-tinged, margins sometimes red-brown, linear or narrowly to broadly lanceolate to narrowly oblong, oblanceolate, or ovate, 1–8.5 cm, not fleshy, margins plane or wavy, involute or flat, (0–)3–5(–7)-lobed, apex acute to rounded or acuminate; lobes ascending, linear to narrowly lanceolate, much narrower than mid blade, apex acute to rounded. |
green, rarely purple-tinged, linear-lanceolate, sometimes linear or lanceolate, 1.3–6.5 cm, not fleshy, margins plane, flat or involute, entire, apex acute. |
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Inflorescences | 3–16(–30 in fruit) × 2–5 cm; bracts proximally greenish to dull reddish purple, distally red to orange or yellow, sometimes crimson, scarlet, orange-red, red-orange, or burnt orange, often becoming paler and/or duller with age, lanceolate to oblong, ovate, or obovate, 3–5(–11)-lobed; lobes spreading to ascending, linear to oblong or narrowly lanceolate, medium length or long, arising at or above mid length, central lobe apex obtuse to rounded or truncate, sometimes emarginate, truncate, or acute, lateral ones acute to obtuse. |
2.5–9 × 1.5–3 cm; bracts red, crimson, scarlet, pink, magenta, red-orange, burnt orange, orange, pale yellow, or whitish throughout, or proximally greenish, distally as stated above, oblong, narrowly obovate, elliptic-oblong, or narrowly ovate, 0(–5)-lobed, rarely with 1 or 2 pairs of short, usually distal lobes; lobes ascending, lanceolate, very short, arising from distal edge, apex rounded to obtuse. |
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Pedicels | 0 mm. |
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Corollas | slightly curved, 17–38 mm; tube 12–18 mm; beak exserted, adaxially green, sometimes yellowish, rarely red-brown, 9–20 mm; abaxial lip ascending, deep green, reduced, curved, 0.5–3 mm, to 10–33% length of beak; teeth incurved, reduced, green, 0.5–1.2 mm. |
straight, 20–33 mm; tubes 13–18 mm; beak, and sometimes abaxial lip, partially to fully exserted; beak adaxially green to yellowish, 8–15 mm; abaxial lip incurved, green, thickened, 2–3 mm, 20–33% as long as beak; teeth ascending or incurved, green, 0.5–1 mm. |
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Calyces | colored as bracts, sometimes with a yellow band proximal to red to orange apices, or ca. 1/2 yellowish and 1/2 reddish, 12–35 mm; abaxial and adaxial clefts 7–12 mm, 33–65% of calyx length, deeper than laterals, lateral 2–7 mm, 15–30% of calyx length; lobes triangular, linear, or oblong to lanceolate, apex acute or obtuse to rounded. |
proximally green to pale green, distally colored as bracts, 15–25 mm; abaxial and adaxial clefts 5–14 mm, 33–50% of calyx length, deeper than laterals, lateral 1–4 mm, 6–20% of calyx length; lobes lanceolate to narrowly triangular, apex rounded to obtuse. |
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2n | = 24, 48, 96. |
= 48. |
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Castilleja hispida |
Castilleja elmeri |
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Phenology | Flowering May–Aug. | |||||
Habitat | Moist to dry rocky slopes, meadows, swales, sagebrush steppes, open conifer forests, montane to subalpine, often on serpentine. | |||||
Elevation | 600–2600 m. (2000–8500 ft.) | |||||
Distribution |
ID; MT; OR; WA; AB; BC
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WA; BC
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Discussion | Varieties 2 (2 in the flora). Castilleja hispida is likely related to C. chromosa, which replaces it geographically to the southeast. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The bracts and inflorescences of Castilleja elmeri can be red, crimson, scarlet, pink, magenta, red-orange, burnt orange, orange, pale yellow, or whitish. While the coloration is highly variable between local populations, it is usually uniform and consistent within them. It is found primarily in the Wenatchee Mountains of Washington, where it is often on serpentine, but it ranges northward in the Cascades into extreme southern British Columbia, apparently on non-serpentine substrates. Hybrids with C. miniata var. miniata, C. parviflora var. albida, and C. thompsonii have been found in the eastern Cascades of Washington, and a similar hybrid swarm with C. thompsonii is reported from southern British Columbia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 612. | FNA vol. 17, p. 605. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Bentham: in W. J. Hooker, Fl. Bor.-Amer. 2: 105. (1838) | Fernald: Erythea 6: 51. (1898) — (as Castilleia) | ||||
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