Castilleja glandulifera |
Castilleja lutescens |
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gland Indian paintbrush, glandular paintbrush, sticky paintbrush |
stiff yellow Indian paintbrush, stiff yellow paintbrush, yellow paintbrush, yellowish paintbrush |
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Habit | Herbs, perennial, 1–3 dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.4–6 dm; from a woody caudex; with a taproot. |
Stems | few to many, erect or ascending, sometimes decumbent, unbranched or often branched proximally, hairs spreading, medium length and long, soft, mixed with more abundant stipitate-glandular ones. |
few to several, erect or ascending, sometimes decumbent at base, unbranched or branched, hairs spreading to retrorse, short, scabrid below inflorescence, sometimes becoming medium length to long, soft to stiff, mixed with short stipitate-glandular ones in inflorescence. |
Leaves | green, linear-lanceolate to sometimes narrowly oblong or narrowly oblanceolate, 0.7–3.7 cm, not fleshy, margins wavy, involute, 0(–5)-lobed, apex acute; lateral lobes ascending to erect, narrowly lanceolate to narrowly oblong, usually narrower than center lobe, apex acute. |
green, sometimes purplish, linear to lanceolate or narrowly oblong, 1–8.5 cm, not fleshy, margins plane, flat or involute lengthwise, 0(–5)-lobed, apex acute; lobes ascending, linear, central one sometimes shallowly toothed, apex acute to obtuse. |
Inflorescences | 2.5–10 × 2–5 cm; bracts proximally pale green to pale yellow, distally yellow, whitish, pink, dull red, or purple on apices (sometimes gradually differentiated from proximal coloration), lanceolate, broadly lanceolate, or oblong, 3–5(–7)-lobed, sometimes with secondary lobes; lobes ascending to spreading, linear, sometimes rounded, medium length or distal short, arising near mid length, apex acute to rarely obtuse. |
3–14 × 1.5–3 cm; bracts greenish throughout, or proximally greenish, distally pale to bright yellow or whitish, rarely pale orangish, lanceolate to oblong, (0–)3–7-lobed; lobes ascending to erect, linear to lanceolate or narrowly oblong, medium length, arising at or above mid length, central lobe apex obtuse to rounded, lateral ones acute. |
Corollas | straight or slightly curved, (20–)22–30 mm; tube 15 mm; abaxial lip usually hidden or just visible in abaxial calyx notch, not exserted/longer than calyx, beak exserted; beak straight or slightly curved, adaxially green, 8–11(–12) mm; abaxial lip deep green to yellow, reduced, slightly pouched, 1–2.5 mm, to 20% as long as beak; teeth incurved, green to yellow, 0.5–1 mm. |
straight or slightly curved, 21–27 mm; tube 14–16 mm; beak partly exserted, adaxially green, 7–12 mm; abaxial lip ascending, green, reduced, 2–4 mm, 25–50% as long as beak; teeth erect or incurved, sometimes spreading, green or white, 0.7–2.5 mm. |
Calyces | proximally green or pale, distally colored as bracts, 17–21(–23) mm; abaxial and adaxial clefts 4–8 mm, 33–50% of calyx length, deeper than laterals, lateral 2–6 mm, 15–33% of calyx length; lobes linear, narrowly lanceolate, or narrowly triangular to oblong, apex acute. |
colored as bracts, 15–25 mm; abaxial and adaxial clefts 6–13 mm, 50% of calyx length, deeper than laterals, lateral 1–7 mm, 15% of calyx length; lobes narrowly triangular to linear, apex acute. |
2n | = 48, 96. |
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Castilleja glandulifera |
Castilleja lutescens |
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Phenology | Flowering Jun–Aug. | Flowering May–Aug. |
Habitat | Dry sagebrush steppes, gravelly or rocky slopes, talus, open conifer forests, subalpine. | Grasslands, open conifer forests, moist meadows, rocky slopes, valleys, montane. |
Elevation | 1400–2500 m. (4600–8200 ft.) | 600–1900 m. (2000–6200 ft.) |
Distribution |
OR
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ID; MT; OR; WA; AB; BC
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Discussion | Castilleja glandulifera is endemic to the upper elevations of the Blue and Strawberry mountains of northeastern Oregon, as well as a few adjacent minor ranges. It is related to C. applegatei and C. viscidula, which are the source of reports of C. glandulifera in the Wallowa Mountains and on Steens Mountain. Inflorescences of C. glandulifera are usually white to pale yellow, but in the area around Marble Creek Pass in Baker County, they are multicolored, with a variety of reddish shades mixed in among the yellowish plants. Castilleja glandulifera and C. viscidula share a glandular pubescence, divided leaves, and usually yellowish inflorescences. Castilleja glandulifera is distinguished from C. viscidula by its taller stature, longer corolla beak, and more deeply divided leaves and bracts with linear to linear-lanceolate lobes. Castilleja glandulifera differs from C. applegatei by its unusual leaves and bracts as well as by its habitat and narrower and somewhat shorter corolla beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja lutescens is found east of the Cascade Range in Oregon and Washington and ranges east to western Montana and the adjacent interior of western Canada. Its inflorescences vary in color from entirely greenish to white or yellowish, and the stature of the plants is also variable, trending from moderate and compact in grasslands to taller in more forested situations. In the Blue Mountains of Garfield County, Washington, occasional hybrids form between C. lutescens and C. hispida var. acuta, which often both occur in the same vicinity. L. R. Heckard (1968) reported a chromosome count of ca. 2n = 120 from a population in Montana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 610. | FNA vol. 17, p. 624. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. pallida var. lutescens | |
Name authority | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 8. (1941) | (Greenman) Rydberg: Mem. New York Bot. Gard. 1: 359. (1900) |
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