Castilleja glandulifera |
Castilleja cusickii |
|
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gland Indian paintbrush, glandular paintbrush, sticky paintbrush |
Cusick's Indian paintbrush, Cusick's paintbrush |
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Habit | Herbs, perennial, 1–3 dm; from a woody caudex; with a taproot. | Herbs, perennial, (1–)1.5–5(–6) dm; from a small, woody caudex; with a taproot or stout, branched roots. |
Stems | few to many, erect or ascending, sometimes decumbent, unbranched or often branched proximally, hairs spreading, medium length and long, soft, mixed with more abundant stipitate-glandular ones. |
solitary or few to several, erect, often decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, eglandular, mixed with shorter, sometimes stiff, stipitate-glandular ones. |
Leaves | green, linear-lanceolate to sometimes narrowly oblong or narrowly oblanceolate, 0.7–3.7 cm, not fleshy, margins wavy, involute, 0(–5)-lobed, apex acute; lateral lobes ascending to erect, narrowly lanceolate to narrowly oblong, usually narrower than center lobe, apex acute. |
green, sometimes with prominent red-purple veins, lanceolate-linear to broadly lanceolate, 2.5–4.5(–7) cm, not fleshy, margins plane, flat, (0–)3–5(–9)-lobed, apex acuminate to acute; lobes spreading-ascending, narrowly lanceolate, arising at or above mid length, apex acute. |
Inflorescences | 2.5–10 × 2–5 cm; bracts proximally pale green to pale yellow, distally yellow, whitish, pink, dull red, or purple on apices (sometimes gradually differentiated from proximal coloration), lanceolate, broadly lanceolate, or oblong, 3–5(–7)-lobed, sometimes with secondary lobes; lobes ascending to spreading, linear, sometimes rounded, medium length or distal short, arising near mid length, apex acute to rarely obtuse. |
3.5–26 × 1.5–3.5 cm; bracts pale green, pale greenish yellow, or pale yellow throughout, or proximally pale green, pale greenish yellow, pale yellow, or reddish purple, distally white, yellow, pink, dull purple, or dull reddish purple, sometimes with a purple band below that, sometimes with veins darker than background color, lanceolate to oblong, 0–5(–9)-lobed; lobes ascending or spreading, linear to narrowly lanceolate, often short, arising above mid length, central lobe apex rounded, lateral ones acute to rounded. |
Corollas | straight or slightly curved, (20–)22–30 mm; tube 15 mm; abaxial lip usually hidden or just visible in abaxial calyx notch, not exserted/longer than calyx, beak exserted; beak straight or slightly curved, adaxially green, 8–11(–12) mm; abaxial lip deep green to yellow, reduced, slightly pouched, 1–2.5 mm, to 20% as long as beak; teeth incurved, green to yellow, 0.5–1 mm. |
straight, 19–25(–28) mm; tube (13–)15–20 mm; abaxial lip included, beak not or slightly exserted; beak adaxially green to yellow-green, 4–7 mm; abaxial lip yellow, prominent, inflated, 2–4.5 mm, ca. 75% as long as beak; teeth incurved to erect, white, 1.3–2.1 mm. |
Calyces | proximally green or pale, distally colored as bracts, 17–21(–23) mm; abaxial and adaxial clefts 4–8 mm, 33–50% of calyx length, deeper than laterals, lateral 2–6 mm, 15–33% of calyx length; lobes linear, narrowly lanceolate, or narrowly triangular to oblong, apex acute. |
green, pale green, or pale yellow, lobes yellow to pale yellow, 20–30 mm; abaxial and adaxial clefts 6.6–14 mm, 33–50% of calyx length, deeper than laterals, lateral 0–4(–4.5) mm, 0–15% of calyx length; lobes broadly rounded to triangular, apex obtuse to rounded. |
2n | = 24. |
|
Castilleja glandulifera |
Castilleja cusickii |
|
Phenology | Flowering Jun–Aug. | Flowering Apr–Aug. |
Habitat | Dry sagebrush steppes, gravelly or rocky slopes, talus, open conifer forests, subalpine. | Moist meadows, swales, grasslands in sagebrush steppes, occasionally to subalpine. |
Elevation | 1400–2500 m. (4600–8200 ft.) | 500–2500 m. (1600–8200 ft.) |
Distribution |
OR
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ID; MT; NV; OR; WA; WY; AB; BC
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Discussion | Castilleja glandulifera is endemic to the upper elevations of the Blue and Strawberry mountains of northeastern Oregon, as well as a few adjacent minor ranges. It is related to C. applegatei and C. viscidula, which are the source of reports of C. glandulifera in the Wallowa Mountains and on Steens Mountain. Inflorescences of C. glandulifera are usually white to pale yellow, but in the area around Marble Creek Pass in Baker County, they are multicolored, with a variety of reddish shades mixed in among the yellowish plants. Castilleja glandulifera and C. viscidula share a glandular pubescence, divided leaves, and usually yellowish inflorescences. Castilleja glandulifera is distinguished from C. viscidula by its taller stature, longer corolla beak, and more deeply divided leaves and bracts with linear to linear-lanceolate lobes. Castilleja glandulifera differs from C. applegatei by its unusual leaves and bracts as well as by its habitat and narrower and somewhat shorter corolla beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja cusickii includes two morphologically distinct variants. One variant has a wider inflorescence, broader bracts, and wider bract lobes. These bracts often have highly contrasting purple veins, and some populations also have a purplish wash on the bracts. These plants have a range on the western edge of typical C. cusickii, though overlapping with more typical forms in western Idaho and adjacent northeastern Oregon and southeastern Washington. The name C. lutea is available for these variants, and both consistent morphological differences and preliminary results from ongoing genetic research on the plants (D. C. Tank, pers. comm.) indicate its resurrection may be justified. A second variant is distinguished by a pale pink-purple wash on the bracts and a narrow distribution in the meadows of the Bear River Valley, bordering northwestern Utah and adjacent southeastern Idaho. Only bract coloration appears to distinguish this form from typical C. cusickii, and the distinctive bract coloration is only more or less consistent in these populations. Typical Castilleja cusickii appears to hybridize with C. gracillima in the Logan Valley, Grant County, Oregon. It is also reported by M. Ownbey (1959) to hybridize with C. miniata and C. rhexiifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 610. | FNA vol. 17, p. 600. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. lutea | |
Name authority | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 8. (1941) | Greenman: Bot. Gaz. 25: 267. (1898) — (as Castilleia) |
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