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wavyleaf Indian paintbrush

Heckard's Indian paintbrush, Heckard's paintbrush

Habit Herbs, perennial, 1.4–8 dm; from a woody caudex; with a taproot. Herbs or subshrubs, perennial, 1.5–4.5 dm; from a woody caudex; with a taproot.
Stems

many, erect, unbranched or branched, sometimes with small, leafy axillary branches, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones.

few to several, decumbent to erect, sometimes leaning, unbranched or often much-branched distally, with a few short, leafy axillary shoots, hairs spreading, short, soft, stipitate-glandular, mixed with long-spreading, eglandular ones.

Leaves

green or purple-tinged, lanceolate to linear-oblong, 0.8–6.1 cm, not fleshy, margins wavy, involute, 0(–3)-lobed, apex rounded to acute;

lobes spreading-ascending, narrowly lanceolate, apex acute to acuminate.

gray-green, sometimes green, lanceolate-linear to narrowly lanceolate, 1–6.5 cm, not fleshy, margins plane, sometimes wavy, flat to involute, 0(–3)-lobed, apex acuminate;

lobes spreading-ascending, linear to narrowly lanceolate, apex acute.

Inflorescences

3–34 × 1.5–3.5 cm;

flowers usually distichous, remote, except distalmost;

bracts proximally greenish, distally greenish, reddish, orange-red, or dull orange, rarely yellow, proximal bracts frequently greenish throughout, lanceolate to narrowly ovate or narrowly oblong-elliptic, 0(–3)-lobed, proximal wavy-margined;

lobes ascending, triangular, short, arising near apex, apex acute to obtuse.

3–30 × 3–4 cm;

bracts proximally green to dark purplish, distally red to crimson, sometimes pale salmon, linear-lanceolate to broadly lanceolate, 3–5-lobed;

lobes spreading, linear, long, arising below mid length, apex acute, sometimes obtuse.

Corollas

± curved, 25–38 mm;

tube 14–18 mm;

beak and often abaxial lip exserted;

beak adaxially dull orange or dull red, rarely yellow, 14–19 mm;

abaxial lip slightly curved, green, red, whitish, or yellow, not inflated, 2–2.5 mm, 15% as long as beak;

teeth incurved, deep green to reddish, 1–1.5 mm.

straight or slightly curved, 20–40 mm;

tube 15–23 mm;

abaxial lip exserted to included, beak much exserted;

beak adaxially yellow-green to reddish, 9–18 mm;

abaxial lip green, reduced, 0.5–1.5 mm, 5–20% as long as beak;

teeth incurved, green, (0–)0.5–1.5 mm.

Calyces

proximally whitish, green, or purple, distally colored as bracts, 9–18 mm;

abaxial clefts 6–6.5 mm, adaxial 7–9 mm, clefts 33+% of calyx length, deeper than laterals, lateral 1.5–4 mm, ca. 10% of calyx length;

lobes linear-lanceolate, apex acute.

colored as bracts, 15–20 mm;

abaxial clefts 3.4–6.2 mm, adaxial 4.5–9 mm, clefts 25–33% of calyx length, deeper than laterals, lateral 0.5–2 mm, 5–10% of calyx length;

lobes narrowly triangular, often slightly unequal, apex acute.

2n

= 24.

= 48, 72.

Castilleja disticha

Castilleja montigena

Phenology Flowering (May–)Jun–Aug(–Sep). Flowering May–Aug.
Habitat Open conifer forests, rocky or sandy slopes, montane to subalpine. Dry rocky slopes, ledges, open conifer forests, thickets, washes.
Elevation 1600–3000 m. (5200–9800 ft.) 1900–2900 m. (6200–9500 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Castilleja disticha is limited to the central and southern Sierra Nevada. Although similar to C. applegatei and C. martini, C. disticha is distinctive and unique in its long, colorful, and highly exserted corollas with the beak exceeding the tube in length, calyces more brightly colored than the often unlobed bracts, and tall, strongly distichous growth form.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Castilleja montigena is endemic to the northeastern portion of the San Bernardino Mountains of southern California. In the field, this species is consistently and relatively easily distinguished from nearby populations of C. martini var. martini, which it essentially replaces in the northeastern portion of the San Bernardino Mountains. It is apparently of allopolyploid hybrid origin between C. martini var. martini and C. chromosa, which approaches its range from the adjacent Mojave Desert.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 603. FNA vol. 17, p. 632.
Parent taxa Orobanchaceae > Castilleja Orobanchaceae > Castilleja
Sibling taxa
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lanata, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. martini, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. montigena, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurascens, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. wootonii, C. xanthotricha
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. disticha, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lanata, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. martini, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurascens, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. wootonii, C. xanthotricha
Synonyms C. applegatei subsp. disticha
Name authority Eastwood: Proc. Calif. Acad. Sci., ser. 3, 2: 289. (1902) — (as Castilleia) Heckard: Syst. Bot. 5: 83, fig. 17 [center]. (1980)
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