Castilleja disticha |
Castilleja glandulifera |
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wavyleaf Indian paintbrush |
gland Indian paintbrush, glandular paintbrush, sticky paintbrush |
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Habit | Herbs, perennial, 1.4–8 dm; from a woody caudex; with a taproot. | Herbs, perennial, 1–3 dm; from a woody caudex; with a taproot. |
Stems | many, erect, unbranched or branched, sometimes with small, leafy axillary branches, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones. |
few to many, erect or ascending, sometimes decumbent, unbranched or often branched proximally, hairs spreading, medium length and long, soft, mixed with more abundant stipitate-glandular ones. |
Leaves | green or purple-tinged, lanceolate to linear-oblong, 0.8–6.1 cm, not fleshy, margins wavy, involute, 0(–3)-lobed, apex rounded to acute; lobes spreading-ascending, narrowly lanceolate, apex acute to acuminate. |
green, linear-lanceolate to sometimes narrowly oblong or narrowly oblanceolate, 0.7–3.7 cm, not fleshy, margins wavy, involute, 0(–5)-lobed, apex acute; lateral lobes ascending to erect, narrowly lanceolate to narrowly oblong, usually narrower than center lobe, apex acute. |
Inflorescences | 3–34 × 1.5–3.5 cm; flowers usually distichous, remote, except distalmost; bracts proximally greenish, distally greenish, reddish, orange-red, or dull orange, rarely yellow, proximal bracts frequently greenish throughout, lanceolate to narrowly ovate or narrowly oblong-elliptic, 0(–3)-lobed, proximal wavy-margined; lobes ascending, triangular, short, arising near apex, apex acute to obtuse. |
2.5–10 × 2–5 cm; bracts proximally pale green to pale yellow, distally yellow, whitish, pink, dull red, or purple on apices (sometimes gradually differentiated from proximal coloration), lanceolate, broadly lanceolate, or oblong, 3–5(–7)-lobed, sometimes with secondary lobes; lobes ascending to spreading, linear, sometimes rounded, medium length or distal short, arising near mid length, apex acute to rarely obtuse. |
Corollas | ± curved, 25–38 mm; tube 14–18 mm; beak and often abaxial lip exserted; beak adaxially dull orange or dull red, rarely yellow, 14–19 mm; abaxial lip slightly curved, green, red, whitish, or yellow, not inflated, 2–2.5 mm, 15% as long as beak; teeth incurved, deep green to reddish, 1–1.5 mm. |
straight or slightly curved, (20–)22–30 mm; tube 15 mm; abaxial lip usually hidden or just visible in abaxial calyx notch, not exserted/longer than calyx, beak exserted; beak straight or slightly curved, adaxially green, 8–11(–12) mm; abaxial lip deep green to yellow, reduced, slightly pouched, 1–2.5 mm, to 20% as long as beak; teeth incurved, green to yellow, 0.5–1 mm. |
Calyces | proximally whitish, green, or purple, distally colored as bracts, 9–18 mm; abaxial clefts 6–6.5 mm, adaxial 7–9 mm, clefts 33+% of calyx length, deeper than laterals, lateral 1.5–4 mm, ca. 10% of calyx length; lobes linear-lanceolate, apex acute. |
proximally green or pale, distally colored as bracts, 17–21(–23) mm; abaxial and adaxial clefts 4–8 mm, 33–50% of calyx length, deeper than laterals, lateral 2–6 mm, 15–33% of calyx length; lobes linear, narrowly lanceolate, or narrowly triangular to oblong, apex acute. |
2n | = 24. |
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Castilleja disticha |
Castilleja glandulifera |
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Phenology | Flowering (May–)Jun–Aug(–Sep). | Flowering Jun–Aug. |
Habitat | Open conifer forests, rocky or sandy slopes, montane to subalpine. | Dry sagebrush steppes, gravelly or rocky slopes, talus, open conifer forests, subalpine. |
Elevation | 1600–3000 m. (5200–9800 ft.) | 1400–2500 m. (4600–8200 ft.) |
Distribution |
CA
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OR
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Discussion | Castilleja disticha is limited to the central and southern Sierra Nevada. Although similar to C. applegatei and C. martini, C. disticha is distinctive and unique in its long, colorful, and highly exserted corollas with the beak exceeding the tube in length, calyces more brightly colored than the often unlobed bracts, and tall, strongly distichous growth form. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja glandulifera is endemic to the upper elevations of the Blue and Strawberry mountains of northeastern Oregon, as well as a few adjacent minor ranges. It is related to C. applegatei and C. viscidula, which are the source of reports of C. glandulifera in the Wallowa Mountains and on Steens Mountain. Inflorescences of C. glandulifera are usually white to pale yellow, but in the area around Marble Creek Pass in Baker County, they are multicolored, with a variety of reddish shades mixed in among the yellowish plants. Castilleja glandulifera and C. viscidula share a glandular pubescence, divided leaves, and usually yellowish inflorescences. Castilleja glandulifera is distinguished from C. viscidula by its taller stature, longer corolla beak, and more deeply divided leaves and bracts with linear to linear-lanceolate lobes. Castilleja glandulifera differs from C. applegatei by its unusual leaves and bracts as well as by its habitat and narrower and somewhat shorter corolla beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 603. | FNA vol. 17, p. 610. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. applegatei subsp. disticha | |
Name authority | Eastwood: Proc. Calif. Acad. Sci., ser. 3, 2: 289. (1902) — (as Castilleia) | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 8. (1941) |
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