Castilleja disticha |
Castilleja campestris |
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wavyleaf Indian paintbrush |
field owl's-clover, meadow paintbrush, vernal pool Indian paintbrush, yellow owl's clover |
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Habit | Herbs, perennial, 1.4–8 dm; from a woody caudex; with a taproot. | Herbs, annual, (0.3–)0.9–3 dm; with fibrous roots. | ||||
Stems | many, erect, unbranched or branched, sometimes with small, leafy axillary branches, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones. |
solitary or few, erect, unbranched, sometimes branched, glabrous or hairs sparse proximally, spreading, long, soft distally. |
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Leaves | green or purple-tinged, lanceolate to linear-oblong, 0.8–6.1 cm, not fleshy, margins wavy, involute, 0(–3)-lobed, apex rounded to acute; lobes spreading-ascending, narrowly lanceolate, apex acute to acuminate. |
green, sometimes purple-tinged, linear to narrowly lanceolate, (0.4–)1–4 cm, not fleshy, thin and flexible or thick and ± brittle, margins plane, involute, 0-lobed, apex acuminate. |
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Inflorescences | 3–34 × 1.5–3.5 cm; flowers usually distichous, remote, except distalmost; bracts proximally greenish, distally greenish, reddish, orange-red, or dull orange, rarely yellow, proximal bracts frequently greenish throughout, lanceolate to narrowly ovate or narrowly oblong-elliptic, 0(–3)-lobed, proximal wavy-margined; lobes ascending, triangular, short, arising near apex, apex acute to obtuse. |
2–15 × 2–3 cm; bracts green throughout, sometimes purplish tinged, linear to narrowly lanceolate, 0-lobed, apex acute. |
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Corollas | ± curved, 25–38 mm; tube 14–18 mm; beak and often abaxial lip exserted; beak adaxially dull orange or dull red, rarely yellow, 14–19 mm; abaxial lip slightly curved, green, red, whitish, or yellow, not inflated, 2–2.5 mm, 15% as long as beak; teeth incurved, deep green to reddish, 1–1.5 mm. |
straight, 11–26 mm; tube 10–22 mm; abaxial lip, beak, and part of tube exserted from calyx; beak adaxially white or yellow, 3–6 mm; abaxial lip yellow, sometimes orange, paler near teeth, prominent, deeply saccate with 3 lobes at ca. 90° from one another, pouches 6–10 mm wide, 3–4 mm deep, abruptly widening from tube, sacs ± round or ± rounded-triangular from above, villous within at base of teeth, 4–5 mm (2–4 mm from sinus), 70–90% as long as beak, puberulent; teeth erect, white, sometimes with deeply colored bases, 0.5–2 mm. |
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Calyces | proximally whitish, green, or purple, distally colored as bracts, 9–18 mm; abaxial clefts 6–6.5 mm, adaxial 7–9 mm, clefts 33+% of calyx length, deeper than laterals, lateral 1.5–4 mm, ca. 10% of calyx length; lobes linear-lanceolate, apex acute. |
proximally green, pale, or purplish, distally purple or green, 5.5–11 mm; abaxial and adaxial clefts 2–5 mm, 30–50% of calyx length, deeper than laterals, lateral 1.5–2.5 mm, 25–30% of calyx length; lobes linear-lanceolate to narrowly triangular, apex acute to acuminate. |
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2n | = 24. |
= 24. |
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Castilleja disticha |
Castilleja campestris |
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Phenology | Flowering (May–)Jun–Aug(–Sep). | |||||
Habitat | Open conifer forests, rocky or sandy slopes, montane to subalpine. | |||||
Elevation | 1600–3000 m. (5200–9800 ft.) | |||||
Distribution |
CA
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CA; OR
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Discussion | Castilleja disticha is limited to the central and southern Sierra Nevada. Although similar to C. applegatei and C. martini, C. disticha is distinctive and unique in its long, colorful, and highly exserted corollas with the beak exceeding the tube in length, calyces more brightly colored than the often unlobed bracts, and tall, strongly distichous growth form. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Castilleja campestris is sometimes confused with Triphysaria eriantha subsp. eriantha, but C. campestris has two-celled anthers and entire leaves, while T. eriantha has one pollen sac per stamen and strongly divided leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 603. | FNA vol. 17, p. 592. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | C. applegatei subsp. disticha | Orthocarpus campestris | ||||
Name authority | Eastwood: Proc. Calif. Acad. Sci., ser. 3, 2: 289. (1902) — (as Castilleia) | (Bentham) T. I. Chuang & Heckard: Syst. Bot. 16: 656. (1991) | ||||
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