Castilleja disticha |
Castilleja brevistyla |
|
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wavyleaf Indian paintbrush |
short-style owl's-clover, shortstyle Indian paintbrush |
|
Habit | Herbs, perennial, 1.4–8 dm; from a woody caudex; with a taproot. | Herbs, annual, (0.6–)1–4.3 dm; with fibrous roots. |
Stems | many, erect, unbranched or branched, sometimes with small, leafy axillary branches, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones. |
solitary or few, erect, unbranched, sometimes branched, hairs spreading, short and long, soft and stiff, eglandular. |
Leaves | green or purple-tinged, lanceolate to linear-oblong, 0.8–6.1 cm, not fleshy, margins wavy, involute, 0(–3)-lobed, apex rounded to acute; lobes spreading-ascending, narrowly lanceolate, apex acute to acuminate. |
green to purplish, linear to linear-lanceolate, (0.8–)2–6(–8.7) cm, not fleshy, margins plane, flat, 3–5-lobed, apex acuminate; lobes ascending, linear to narrowly lanceolate, long, apex acuminate to rounded. |
Inflorescences | 3–34 × 1.5–3.5 cm; flowers usually distichous, remote, except distalmost; bracts proximally greenish, distally greenish, reddish, orange-red, or dull orange, rarely yellow, proximal bracts frequently greenish throughout, lanceolate to narrowly ovate or narrowly oblong-elliptic, 0(–3)-lobed, proximal wavy-margined; lobes ascending, triangular, short, arising near apex, apex acute to obtuse. |
5–25 (longer in fruit) × 1–2 cm; bracts proximally greenish to dull reddish brown, distally pink, lavender, magenta, purple-red, or white on apices, narrowly lanceolate, (3–)5-lobed; lobes ascending, linear to narrowly lanceolate with slightly widened apices, medium length to long, arising near or below mid length, apex acute. |
Corollas | ± curved, 25–38 mm; tube 14–18 mm; beak and often abaxial lip exserted; beak adaxially dull orange or dull red, rarely yellow, 14–19 mm; abaxial lip slightly curved, green, red, whitish, or yellow, not inflated, 2–2.5 mm, 15% as long as beak; teeth incurved, deep green to reddish, 1–1.5 mm. |
straight, 15–30 mm; tube 14–23 mm, not expanded distally, majority of it exserted from calyx; beak straight, adaxially white or pink (drying purple), 4–6 mm, pubescent; abaxial lip ± inconspicuous, exserted, pouches 3, 2 mm wide, 1–1.5 mm deep, 3–5 mm, 50–70% as long as beak, white, yellow, or pink with large deep purple, red, or brown spot on each pouch at or extending below middle; teeth erect, white, yellow, or pink, 1–1.5 mm. |
Calyces | proximally whitish, green, or purple, distally colored as bracts, 9–18 mm; abaxial clefts 6–6.5 mm, adaxial 7–9 mm, clefts 33+% of calyx length, deeper than laterals, lateral 1.5–4 mm, ca. 10% of calyx length; lobes linear-lanceolate, apex acute. |
colored as bracts, sometimes proximally yellow, 15–20 mm; abaxial and adaxial clefts 7–9.5 mm, abaxial ca. 33% of calyx length, adaxial ca. 66% of calyx length, deeper than laterals, lateral 3–5.5 mm, 33% of calyx length; lobes ± linear, slender, all 4 similar, apex acute. |
Filaments | glabrous. |
|
2n | = 24. |
= 48. |
Castilleja disticha |
Castilleja brevistyla |
|
Phenology | Flowering (May–)Jun–Aug(–Sep). | Flowering Mar–Jun. |
Habitat | Open conifer forests, rocky or sandy slopes, montane to subalpine. | Arid grasslands in hilly country, sagebrush or alkaline flats. |
Elevation | 1600–3000 m. (5200–9800 ft.) | 50–1200 m. (200–3900 ft.) |
Distribution |
CA
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CA
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Discussion | Castilleja disticha is limited to the central and southern Sierra Nevada. Although similar to C. applegatei and C. martini, C. disticha is distinctive and unique in its long, colorful, and highly exserted corollas with the beak exceeding the tube in length, calyces more brightly colored than the often unlobed bracts, and tall, strongly distichous growth form. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja brevistyla is endemic to the foothills of the southern Sierra Nevada and the southern reaches of the Inner South Coast Ranges, in Kern and adjacent counties. It often grows with other annuals, including C. attenuata, C. densiflora, and C. exserta. The similar C. attenuata has three-lobed bracts and leaves, while C. brevistyla has mostly five-lobed parts, and also differs in corolla structure and spotting. Hybrids between C. brevistyla and C. exserta are known from Kern County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 603. | FNA vol. 17, p. 591. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. applegatei subsp. disticha | Orthocarpus brevistylus |
Name authority | Eastwood: Proc. Calif. Acad. Sci., ser. 3, 2: 289. (1902) — (as Castilleia) | (Hoover) T. I. Chuang & Heckard: Syst. Bot. 16: 656. (1991) |
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