Castilleja cryptantha |
Castilleja affinis |
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Mt. Rainier or obscure paintbrush, obscure Indian paintbrush, obscure paintbrush |
California paintbrush, coast Indian paintbrush, coast paintbrush, Indian paintbrush |
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Habit | Herbs, perennial, (0.8–)1–1.9 dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. | ||||||||
Stems | few to several, erect or ascending, unbranched, hairs spreading, long, soft, eglandular, mixed with short stipitate-glandular ones. |
few to many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. |
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Leaves | green, often with brown or purple veins, narrowly to broadly lanceolate, 1.5–4 cm, not fleshy, margins plane, ± involute, 0–3-lobed, apex acute to acuminate; lobes spreading-ascending, narrowly lanceolate, apex acute to ± obtuse. |
green or purplish, sometimes red-brown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid blade, apex acute to rounded. |
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Inflorescences | (2.5–)3–6 × 1–2 cm; green to dull brown or dull reddish purple throughout, or proximally green to dull brown or dull reddish purple, distally yellow on apices, broadly lanceolate to ovate, (0–)3-lobed; lobes ascending, narrowly lanceolate, long or short, arising near mid length, apex acute or acuminate. |
3–25(–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–)3–5(–7)-lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. |
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Pedicels | 0 mm or nearly so. |
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Corollas | straight, 14–16 mm; tube 11–14 mm; whole corolla included within calyx; beak adaxially pale yellow, 1–2 mm; abaxial lip deep green, slightly inflated, 4–5 mm, 67% as long as beak; teeth ascending, pale, 1.5–2 mm. |
straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. |
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Calyces | proximally green or pale with green veins, lobes yellow, sometimes becoming deep red with age, 12–15 mm; abaxial and adaxial clefts 3–7 mm, 25–50% of calyx length, deeper than laterals, lateral 1–3(–4) mm, 8–20% of calyx length; lobes triangular, adaxial segments longer than abaxials, apex acute or obtuse. |
proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. |
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2n | = 24. |
= 48, 72, 96. |
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Castilleja cryptantha |
Castilleja affinis |
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Phenology | Flowering Jul–Aug. | |||||||||
Habitat | Mesic to moist flat subalpine meadows and turf, to tree line. | |||||||||
Elevation | 1500–2000 m. (4900–6600 ft.) | |||||||||
Distribution |
WA
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CA; nw Mexico
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Discussion | Castilleja cryptantha is endemic to the vicinity of Mt. Rainier in the Cascade Range, with most populations found within Mt. Rainier National Park. Unlike most species of Castilleja, it is apparently self-pollinating (W. J. Duffield 1972); the small flowers are entirely enclosed within the yellowish calyces, which tend to grow deep reddish as they age. The purplish brown bracts are also unusual in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 600. | FNA vol. 17, p. 582. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Pennell & G. N. Jones: Proc. Biol. Soc. Wash. 50: 208. (1937) | Hooker & Arnott: Bot. Beechey Voy., 154. (1833) | ||||||||
Web links |