Castilleja crista-galli |
Castilleja victoriae |
|
---|---|---|
cock's-comb paintbrush, mountainside Indian paintbrush |
Victoria owl-clover, Victoria paintbrush, Victoria's owl's-clover, Victoria's owl-clover, Victoria's paintbrush |
|
Habit | Herbs, perennial, 1–5 dm; from a woody caudex; with a taproot. | Herbs, annual, 0.2–2 dm; with fibrous roots. |
Stems | few to several, erect or ascending, unbranched or branched, sometimes with short, leafy axillary shoots, hairy, sometimes glabrate proximally, hairs spreading to retrorse, medium length to long, soft, eglandular, often mixed distally with shorter stipitate-glandular ones. |
solitary, erect, unbranched, sometimes branched, hairs spreading, long, soft, mixed with short stipitate-glandular ones. |
Leaves | green, linear to narrowly lanceolate, 2–8 cm, not fleshy, margins plane, involute, 0–5-lobed, apex acute; lateral lobes spreading, linear, apex acuminate. |
usually brownish throughout, sometimes greenish proximally, brownish distally, margins deep brown, lanceolate to narrowly ovate, 0.5–2.7 cm, not fleshy, margins plane, involute, 0–3(–5)-lobed, apex acute; lobes spreading-ascending, linear to lanceolate, apex acute, obtuse, or rounded. |
Inflorescences | 3–6(–11) × 1.5–6.5 cm; bracts proximally greenish, distally red, red-orange, or orange, sometimes yellow or dull salmon, narrowly to broadly lanceolate or oblong, 3–5-lobed; lobes ascending-spreading, linear-lanceolate, long, arising below mid length, central lobe apex rounded to obtuse, lateral ones acute. |
1–5 × 1.5–3 cm; bracts dull reddish brown throughout, or proximally dull greenish, distally dull reddish brown, narrowly ovate, 3–7-lobed; lobes ascending, lanceolate, medium length, arising near or above mid length, apex broadly acute or obtuse. |
Corollas | straight, (25–)30–40(–45) mm; tube 15–20 mm; abaxial lip visible through front cleft, beak long-exserted from calyx; beak adaxially green or yellow-green, 16–21 mm; abaxial lip proximally white or yellow-green, distally green, reduced, usually visible in front cleft, 3 mm, 20% as long as beak; teeth incurved to ascending, green, 1 mm. |
straight, 10–18 mm; tube 9–13 mm; abaxial lip and beak exserted; beak adaxially white, sometimes faintly diffuse purple, 3 mm, inconspicuously puberulent, hairs short stipitate-glandular; abaxial lip yellowish, cream, or soft pale yellow, lacking spots, moderately conspicuous, pouches 3, conspicuous, 1–3 mm deep, central one furrowed, conspicuous, 2–2.5 mm, 60–75% as long as beak; teeth erect, white or cream, 0.5–0.8 mm. |
Calyces | colored as bracts, (20–)25–35 mm; abaxial and adaxial clefts (6–)10–17 mm, 50% of calyx length, deeper than laterals, lateral (1–)3–6(–10) mm, 35% of calyx length; lobes slender, triangular, apex acute. |
colored as bracts, 8–12 mm; abaxial and adaxial clefts 5–6 mm, 50% of calyx length, deeper than laterals, lateral 3–4 mm, 30–40% of calyx length; lobes linear-lanceolate, apex acute. |
Stigmas | included in beak. |
|
2n | = 96. |
= 24. |
Castilleja crista-galli |
Castilleja victoriae |
|
Phenology | Flowering Jun–Aug. | Flowering (Apr–)May–Jul. |
Habitat | Rocky slopes, talus, ridges, dry to moist, open, conifer forests, montane meadows. | Vernal pools, springs, windswept, thin-soiled rocky knolls and benches by sea, gneiss. |
Elevation | 1500–2900 m. (4900–9500 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
ID; MT; WY
|
WA; BC |
Discussion | Castilleja crista-galli is found in the Rocky Mountains of southwestern Montana and northwestern Wyoming. The extent of its distribution into adjacent Idaho is unresolved, in part because it is frequently confused with either C. linariifolia or C. miniata. Castilleja crista-galli appears to be morphologically intermediate between them, leading to speculation that it might be an allopolyploid derivative. A DNA study (S. Matthews and M. Lavin 1998) showed little support for a hybrid origin. Castilleja crista-galli may be separated with some difficulty from the other two species by the presence of at least some short hairs on the stems and the frequently three- to five-parted leaves. Castilleja linariifolia and C. miniata both usually have subglabrous stems and entire leaves, sometimes three-parted distally, near the inflorescence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja victoriae was first collected in 1893 but not described until 2008. It is restricted to the southernmost tip of Vancouver Island, near Oak Bay, and to several small adjacent islands within a 30 km radius in British Columbia and Washington. Its primary habitat is small depressions and vernal pools on gentle gradients within 100 m of the sea, making it particularly vulnerable to development and recreational conflicts. Never common, several historic populations near Victoria were extirpated before the species was recognized. The majority of the global population is found on Trial Island. All known extant populations would be inundated by projected sea level increases and storm surges due to climate change. It should be considered globally endangered. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 599. | FNA vol. 17, p. 663. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Name authority | Rydberg: Mem. New York Bot. Gard. 1: 355. (1900) | Fairbarns & J. M. Egger: Madroño 54: 335, figs. 1, 3 [lower right]. (2008) |
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