Castilleja collegiorum |
Castilleja exserta |
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colleagues paintbrush, collegial paintbrush |
castilleja exserta, escobita, exserted Indian paintbrush, owl's clover, purple owl's-clover, purple painted-cup |
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Habit | Herbs, perennial, 1.1–2.8 dm; from a woody caudex; with a taproot. | Herbs, annual, 0.1–4.5 dm; with fibrous roots. | ||||||||
Stems | few to many, erect or ascending, short-decumbent at base, unbranched, hairs dense, spreading to erect, ± short, soft, usually stipitate-glandular, longer ones sometimes eglandular. |
solitary, erect to ascending, unbranched or diffusely branched from near base, hairs spreading, medium length and long, stiff, mixed with short stipitate-glandular ones. |
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Leaves | pale green to dull reddish maroon, linear to linear-lanceolate, 0.8–3.5 cm, not fleshy, margins plane, slightly involute, 0–3(–5)-lobed, apex acuminate; lateral lobes ascending to spreading, linear-lanceolate, usually arising from distal 1/2 of blade, usually narrower than central lobe, apex acuminate to acute. |
green, sometimes purple to brownish, linear or ovate to orbicular in outline, (0.8–)1–5(–7.7) cm, not fleshy, margins plane, involute, (0–)3–9(–11)-lobed, sometimes with secondary lobing, apex acuminate to rounded or acute; lobes spreading, filiform or linear to narrowly spatulate, apex acute to acuminate or rounded. |
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Inflorescences | 10–40(–80 with age) × 0.5–2.5 cm; bracts pale cream to pale greenish yellow throughout, often partly to entirely suffused with dull reddish purple to maroon, especially proximally, along veins, and with age, sometimes distal apices pale, dullish red, lanceolate to ovate, usually 3-lobed, central lobe sometimes with short teeth; lobes spreading-ascending, linear-lanceolate, short to medium length, arising at or above mid length, apex acute. |
1.5–20 × 2–4 cm; bracts proximally greenish, dark purple, brownish purple, or white, distally pink, lavender, magenta, light purple, or white on lobe apices, lanceolate to elliptic or narrowly ovate, (3–)5(–9)-lobed, often with 2–4 secondary lobes; lobes ascending to spreading, linear to filiform or narrowly spatulate, medium length to long, arising below or above mid length, apex rounded to acute. |
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Corollas | straight, 16.5–25 mm; tube 12–18 mm; beak scarcely exserted, adaxially pale green to yellowish, 3–7 mm; abaxial lip green, not inflated, grooved, 2.5 mm, 33–50% as long as beak; teeth slightly incurved, white, 1 mm. |
straight, 12–33 mm; tube 7–20 mm; beak slightly exserted, hooked near apex, adaxially pink-purple to magenta, rarely white, 5–13 mm, margins colored as bracts, densely villous-hairy; abaxial lip proximally pink, purple, or magenta, rarely yellow or white, with maroon or deep purple distal to that and white to yellow or pink distally, often with purple spots, distal pale color often aging deep pink or deep red, strongly inflated, pouches 3, 3–8 mm wide, 3–4 mm deep, 3–9 mm, 67–80% as long as beak; teeth erect, white, yellow, or purple, often with purple or maroon spots, 0.5–2 mm. |
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Calyces | pale cream to pale greenish yellow, sometimes reddish violet to maroon on distal segments and/or with a thin vertical strip of pale reddish violet to maroon along veins, 11–20 mm; abaxial and adaxial clefts 7–12 mm, 60% of calyx length, deeper than laterals, lateral 0.5–1 mm, 5–10% of calyx length; lobes triangular, apex acute to obtuse. |
colored as bracts, 10–26 mm; abaxial clefts 4–12 mm, adaxial 9–18 mm, abaxial ca. 50% of calyx length, adaxial ca. 67% of calyx length, at least adaxial deeper than others, lateral 2.5–9 mm, 15–45% of calyx length; lobes linear (to narrowly oblanceolate), apex rounded to acute. |
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Filaments | with spreading, long, soft hairs. |
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2n | = 24. |
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Castilleja collegiorum |
Castilleja exserta |
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Phenology | Flowering late Jun–Jul. | |||||||||
Habitat | Hummocks and margins of moist to wet meadows. | |||||||||
Elevation | 1700–1800 m. (5600–5900 ft.) | |||||||||
Distribution |
OR
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AZ; CA; NM; nw Mexico
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Discussion | Castilleja collegiorum is endemic to a large meadow system in the southern Cascade Range of Klamath County. It is similar to C. cryptantha in Washington and C. lemmonii in California but differs from both in structural details of the inflorescence, calyx, bracts, and leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). In addition to the characters in the key, Castilleja exserta is distinguished from the similar C. densiflora by its conspicuously hairy and apically hooked beak. As a result, the capitate stigma is exserted more or less horizontally from the corolla beak. In contrast, C. densiflora has an unhooked, inconspicuously puberulent beak, from which the stigma emerges more vertically. Castilleja exserta hybridizes with C. attenuata in southern California and with C. lineariloba in central California, and it reportedly crosses occasionally with C. densiflora in southern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 598. | FNA vol. 17, p. 606. | ||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||
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Synonyms | Orthocarpus exsertus | |||||||||
Name authority | J. M. Egger & S. Malaby: Phytoneuron 2015-33: 1, figs. 1–3, 9[left]. (2015) | (A. Heller) T. I. Chuang & Heckard: Syst. Bot. 16: 657. (1991) | ||||||||
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