Castilleja chrysantha |
Castilleja peckiana |
|
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Wallowa paintbrush, yellow paintbrush, yellow Wallowa Indian paintbrush, yellowish or common Wallowa paintbrush |
peak paintbrush, Peck's Indian paintbrush, Peck's paintbrush |
|
Habit | Herbs, perennial, (0.5–)1–2(–5) dm; from a woody caudex; with a taproot. | Herbs, perennial, (1.8–)2.4–6 dm; from a woody caudex; with a stout taproot. |
Stems | few to many, erect or ascending, often decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, mixed with dense, shorter stipitate-glandular ones. |
few to many, erect or ascending, often branched distally, sometimes unbranched, sometimes with short axillary shoots, proximal hairs retrorse to appressed, short to moderately long, distal hairs spreading, longer, soft, often mixed with short stipitate-glandular ones. |
Leaves | green or purple-tinged, sometimes deep purple, broadly lanceolate, sometimes linear-lanceolate or narrowly oblong, 1.5–4.8 cm, not fleshy, margins plane, flat or involute, 0–3-lobed, apex acute; lobes ascending or spreading, linear to narrowly linear or narrowly lanceolate, short, apex acuminate. |
green, linear-lanceolate, rarely broadly lanceolate, (1.2–)2.5–8(–9) cm, not fleshy, margins plane, flat or involute, 0(–3)-lobed, apex acute to rounded; lobes ascending-spreading, narrowly lanceolate to linear, apex acute or obtuse. |
Inflorescences | 3–17 × 1–3 cm; bracts greenish or pale yellow-green throughout, or proximally greenish or pale yellow-green, distally pale yellow to whitish, sometimes pink-purple, or pale, dull purplish, sometimes aging pink or yellow, often infused with light purple, rarely pink, ovate to broadly acute, (0–)3-lobed; lobes ascending, linear-lanceolate, oblong, or triangular, short, arising below mid length, apex acute to obtuse. |
(2–)4–17 × 1.5–3 cm; bracts proximally greenish, distally red, orange-red, or orange, proximal sometimes lanceolate, distal broadly lanceolate to ovate, (0–)3(–7)-lobed; lobes spreading to ascending, linear to lanceolate, long, arising near or below mid length, central lobe apex rounded, lateral ones acute. |
Corollas | straight, (16–)20–25 mm; tube 15–18 mm; beak slightly exserted, adaxially green to yellow, 5.5–8.5 mm; abaxial lip white, green and white, pink, or purple, ± prominent, appressed (proximally scarcely or not pouched), 3–5 mm, 67% as long as beak; teeth erect, white or pink with some purple or red, 1.5–2.5 mm. |
straight, 23–30(–35) mm; tube 12–20 mm; beak exserted, adaxially green, 8–12(–14) mm; abaxial lip pale to deep green, reduced, rounded, 0.5–1.5 mm, 10–20% as long as beak; teeth erect to incurved, green, (0.4–)0.7–1.2(–2) mm. |
Calyces | proximally green, pale with green veins, purple-tinged green, or purple, distally pale yellow, white, or purplish, 12–20 mm; abaxial and adaxial clefts 8–12 mm, 50% of calyx length, deeper than laterals, lateral 0.5–3 mm, 5–15% of calyx length; lobes short-triangular, apex obtuse, rounded, or truncate. |
proximally pale yellow or greenish, distally colored as bracts, (15–)18–28 mm; abaxial and adaxial clefts 6–12 mm, 40–45% of calyx length, deeper than laterals, lateral 2.5–8 mm, 20–30% of calyx length; lobes linear to narrowly lanceolate, apex acute to acuminate. |
2n | = 72, 96, ca. 120. |
|
Castilleja chrysantha |
Castilleja peckiana |
|
Phenology | Flowering May–Oct. | Flowering (Apr–)May–Aug. |
Habitat | Flat, mesic meadows, dry talus and ridges, montane to alpine. | Open conifer forests, sagebrush slopes, riparian meadows, shores. |
Elevation | 1100–2800 m. (3600–9200 ft.) | 1400–2600 m. (4600–8500 ft.) |
Distribution |
OR
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ID; NV; OR
|
Discussion | Castilleja chrysantha is endemic to the mountains of northeastern Oregon, and its patterns of variation need further study. Most plants in the Blue Mountains are taller, more erect, and tolerate lower elevation habitats than populations around the type locality in the Wallowa Mountains. Plants with purplish inflorescences and longer hairs were described as C. ownbeyana and tend to favor drier talus and ridges than plants like the type, found in flat, mesic, montane to subalpine meadows. Hybrids between C. chrysantha and C. fraterna occur in Wallowa County. In the Wallowa Mountains, a recurrent and variable hybrid form between C. chrysantha and C. rhexiifolia was described as C. wallowensis Pennell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
When describing Castilleja peckiana, Pennell noted that its variation approached C. hispida on one hand and C. miniata on the other, and it is likely of allopolyploid derivation. L. R. Heckard (1968) found chromosome numbers of n = 36, 48, and ca. 60. He hypothesized that C. chromosa, C. hispida var. acuta, and C. miniata were likely involved in its ancestry, and possibly C. pruinosa as well. Heckard suggested subsequent introgression among the derived forms introduced further complexity. Though complex, these forms are self-perpetuating and appear morphologically stable within their range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 596. | FNA vol. 17, p. 643. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. ownbeyana | |
Name authority | Greenman: Bot. Gaz. 48: 146. (1909) | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 9. (1941) |
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