Castilleja chromosa |
Castilleja pallescens |
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desert paintbrush |
dwarf pale paintbrush, pale Indian paintbrush, pale paintbrush |
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Habit | Herbs, sometimes subshrubs, perennial, 1.5–3.5(–4.5) dm; from a woody caudex; with a taproot. | Herbs, perennial, (0.4–)1–3 dm; from a woody caudex; with a taproot. | ||||
Stems | several to many, ascending to erect, often grayish, unbranched, rarely branched, sometimes with short, leafy axillary branches, hairs spreading-erect, long, stiff, eglandular, sometimes also with shorter, stipitate-glandular ones. |
few to many, erect to ascending, decumbent at base, unbranched, sometimes branched, hairs moderately to very dense, retrorsely curved to appressed, short, ± stiff, eglandular. |
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Leaves | gray-green, linear, lanceolate, or oblanceolate, sometimes broadly lanceolate, (1.5–)2.5–6(–7) cm, not fleshy, margins plane, involute, (0–)3–5(–7)-lobed, sometimes with secondary lobes, apex acuminate to obtuse; lobes spreading, linear, apex acuminate. |
purple-tinged or deep purple, sometimes green, linear to narrowly lanceolate, 1–4(–5) cm, not fleshy, margins plane, sometimes ± wavy, ± involute, (0–)3–5(–7)-lobed, apex acute; lobes spreading or ascending-spreading, linear, apex acute. |
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Inflorescences | 2.5–15 (much longer in fruit) × 1.5–5.5 cm; bracts proximally greenish to dull purplish, distally bright red to scarlet or orange-red, rarely yellowish to dull orange or pink, narrowly to broadly linear or lanceolate, narrowly ovate, or oblong-lanceolate, (0–)3–7-lobed, rarely with secondary lobes; lobes spreading, linear to oblong, sometimes oblanceolate, often expanded near tip, long, proximal lobes arising below mid length, apex rounded or obtuse to sometimes acute. |
(1.5–)4–8(–12) × 1.5–5.5 cm; bracts pale green to yellow-green or reddish purple throughout, or proximally pale green to yellow-green, distally white to cream or pale yellowish, sometimes pink to reddish purple, lanceolate to linear-lanceolate, elliptic, or ovate, 3–5(–9)-lobed; lobes spreading to ascending, linear, long, arising along distal 2/3, apex acute to obtuse. |
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Corollas | straight or ± curved, 18–35(–40) mm; tube 8–15 mm; beak short- or long-exserted, adaxially green to yellow-green, (9–)10–18 mm; abaxial lip deep green, reduced, thickened, included to exserted, 2–3 mm, ca. 20% as long as beak; teeth incurved, deep green, 0.5–1 mm. |
straight, 13–23(–27) mm; tube 10–20 mm; subequal to calyx or beak slightly exserted; beak adaxially whitish or buff, rarely pink to pink-purple, 3.5–8 mm; abaxial lip proximally green, white, purple, or purplish brown, distally white, yellow, green, pink, or reddish, prominent, pouched, pouches pleated, longer than deep, gradually expanded, 2.5–8 mm, 70–100% as long as beak, puberulent; teeth erect to spreading, pink, cream, or white, sometimes with a yellow spot proximally, 1.5–3.5 mm. |
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Calyces | colored as bracts, sometimes with broad yellow band below colored lobe apices, (17–)20–27 mm; abaxial clefts 4–10 mm, adaxial 6–12 mm, abaxial ca. 30% of calyx length, adaxial ca. 40% of calyx length, deeper than laterals, lateral 1–4 mm, ca. 15% of calyx length; lobes oblong or ovate to narrowly triangular or lanceolate, apex obtuse to rounded. |
colored as bracts, sometimes distally purple with age, 11–25(–27) mm; abaxial and adaxial clefts 7–13.6 mm, 40–50% of calyx length, deeper than laterals, lateral 0.5–4.3(–6) mm, (0–)5–25% of calyx length; lobes lanceolate to triangular, apex usually triangular or acute, rarely ± obtuse. |
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2n | = 24, 48. |
= 24, 48. |
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Castilleja chromosa |
Castilleja pallescens |
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Phenology | Flowering (Feb–)Mar–Aug(–Nov). | |||||
Habitat | Dry sagebrush slopes and flats, pinyon-juniper stands, blackbrush, open yellow pine forests. | |||||
Elevation | 500–3200 m. (1600–10500 ft.) | |||||
Distribution |
AZ; CA; CO; ID; NM; NV; OR; UT; WY
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ID; MT; NV; OR; WY
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Discussion | Castilleja chromosa is sometimes confused with 3b. C. angustifolia var. dubia (see discussion there). Castilleja chromosa retains its distinctive morphology across its wide range and is a characteristic species of much of the southwestern United States. Where it overlaps with C. angustifolia, the two are distinguished by inflorescence color and width and by the lengths of the calyx, corolla, and corolla beak. In the broad region of their sympatry, there is little evidence of intergradation, except in a few sites in Elko County, Nevada, and in southern Wyoming. Throughout southern Idaho and northeastern Nevada the range of the two overlap with little or no intergradation. At high elevations in Montrose County, Colorado, C. chromosa has narrower leaves and a longer and silkier pubescence, especially in the inflorescence. Apparent hybrids between C. chromosa and C. flava var. rustica are known from Custer County, Idaho, and hybrids with C. linariifolia are known from Montrose County, Colorado. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Castilleja pallescens occurs from valleys to alpine ridges and summits throughout its range, usually in sagebrush communities, but at higher elevations it is also found on dry sites associated with other plant species. The alpine plants are greatly reduced in stature. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 595. | FNA vol. 17, p. 637. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | C. ewanii, C. martini subsp. ewanii, C. martini var. ewanii | Orthocarpus pallescens | ||||
Name authority | A. Nelson: Bull. Torrey Bot. Club 26: 245. (1899) — (as Castilleia) | (A. Gray) Greenman: Bot. Gaz. 25: 266. (1898) — (as Castilleia) | ||||
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