Castilleja chromosa |
Castilleja organorum |
|
---|---|---|
desert paintbrush |
Organ Mountain Indian paintbrush, Organ Mountains paintbrush |
|
Habit | Herbs, sometimes subshrubs, perennial, 1.5–3.5(–4.5) dm; from a woody caudex; with a taproot. | Herbs or subshrubs, perennial, 2.7–8 dm; from a small, woody caudex; with thick, woody roots. |
Stems | several to many, ascending to erect, often grayish, unbranched, rarely branched, sometimes with short, leafy axillary branches, hairs spreading-erect, long, stiff, eglandular, sometimes also with shorter, stipitate-glandular ones. |
several to many, erect to sprawling, usually profusely branched, including many small, leafy axillary shoots, hairs dense, retrorse, medium length, stiff, eglandular, distally spreading, long, soft, sometimes matted, very short-glandular. |
Leaves | gray-green, linear, lanceolate, or oblanceolate, sometimes broadly lanceolate, (1.5–)2.5–6(–7) cm, not fleshy, margins plane, involute, (0–)3–5(–7)-lobed, sometimes with secondary lobes, apex acuminate to obtuse; lobes spreading, linear, apex acuminate. |
green, linear-lanceolate, distally sometimes broadly lanceolate, 2–4.5 cm on main branch, 0.5–1 cm on proximal part of side branches, not fleshy, margins plane, involute, sometimes flat, 0-lobed, apex acute to acuminate. |
Inflorescences | 2.5–15 (much longer in fruit) × 1.5–5.5 cm; bracts proximally greenish to dull purplish, distally bright red to scarlet or orange-red, rarely yellowish to dull orange or pink, narrowly to broadly linear or lanceolate, narrowly ovate, or oblong-lanceolate, (0–)3–7-lobed, rarely with secondary lobes; lobes spreading, linear to oblong, sometimes oblanceolate, often expanded near tip, long, proximal lobes arising below mid length, apex rounded or obtuse to sometimes acute. |
2–4.5(–14 in fruit) × 1.5–4 cm; bracts proximally greenish, distally red to reddish orange, broadly lanceolate to oblong, 0(–3)-lobed; lobes ascending, lanceolate, short, arising above mid length, apex acute to obtuse. |
Corollas | straight or ± curved, 18–35(–40) mm; tube 8–15 mm; beak short- or long-exserted, adaxially green to yellow-green, (9–)10–18 mm; abaxial lip deep green, reduced, thickened, included to exserted, 2–3 mm, ca. 20% as long as beak; teeth incurved, deep green, 0.5–1 mm. |
slightly curved, 15–24 mm; tube 10–13 mm; beak exserted, adaxially green, 6–10 mm; abaxial lip green, reduced, slightly pouched, sometimes visible in front cleft, 0.5–1.5 mm, 15–20% as long as beak; teeth incurved, white or green, 0.4–0.7 mm. |
Calyces | colored as bracts, sometimes with broad yellow band below colored lobe apices, (17–)20–27 mm; abaxial clefts 4–10 mm, adaxial 6–12 mm, abaxial ca. 30% of calyx length, adaxial ca. 40% of calyx length, deeper than laterals, lateral 1–4 mm, ca. 15% of calyx length; lobes oblong or ovate to narrowly triangular or lanceolate, apex obtuse to rounded. |
proximally pale green to pale yellow-green, distally pale red to red-orange above middle, 12.5–20.5 mm; abaxial and adaxial clefts 6–9 mm, 33–50% of calyx length, deeper than laterals, lateral (1.5–)3–4 mm, 20–35% of calyx length; lobes lanceolate or broadly triangular, apex acute. |
2n | = 24, 48. |
|
Castilleja chromosa |
Castilleja organorum |
|
Phenology | Flowering (Feb–)Mar–Aug(–Nov). | Flowering Jun–Oct. |
Habitat | Dry sagebrush slopes and flats, pinyon-juniper stands, blackbrush, open yellow pine forests. | Rocky slopes, shaded canyons, riparian zones, open conifer forests, sun or partial shade. |
Elevation | 500–3200 m. (1600–10500 ft.) | 1500–2500 m. (4900–8200 ft.) |
Distribution |
AZ; CA; CO; ID; NM; NV; OR; UT; WY
|
NM |
Discussion | Castilleja chromosa is sometimes confused with 3b. C. angustifolia var. dubia (see discussion there). Castilleja chromosa retains its distinctive morphology across its wide range and is a characteristic species of much of the southwestern United States. Where it overlaps with C. angustifolia, the two are distinguished by inflorescence color and width and by the lengths of the calyx, corolla, and corolla beak. In the broad region of their sympatry, there is little evidence of intergradation, except in a few sites in Elko County, Nevada, and in southern Wyoming. Throughout southern Idaho and northeastern Nevada the range of the two overlap with little or no intergradation. At high elevations in Montrose County, Colorado, C. chromosa has narrower leaves and a longer and silkier pubescence, especially in the inflorescence. Apparent hybrids between C. chromosa and C. flava var. rustica are known from Custer County, Idaho, and hybrids with C. linariifolia are known from Montrose County, Colorado. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja organorum is endemic to the Organ Mountains of Doña Ana County. Reports from the mountains of the Mogollon Rim of east-central Arizona and adjacent New Mexico are based on specimens of C. nelsonii. Castilleja organorum is grouped with C. linariifolia by some authors (for example, G. L. Nesom 1992c), but it has subequal abaxial and adaxial calyx clefts and is more likely closely related to C. integra. Castilleja organorum differs from the latter species in its loose, often profusely branched habit, more compact inflorescences, and usually smaller corollas. In Fillmore Canyon, C. integra, C. lanata, and C. organorum are all found, but each is in a different habitat, and there is no sign of hybridization. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 595. | FNA vol. 17, p. 636. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. ewanii, C. martini subsp. ewanii, C. martini var. ewanii | |
Name authority | A. Nelson: Bull. Torrey Bot. Club 26: 245. (1899) — (as Castilleia) | Standley: Muhlenbergia 5: 86. (1909) |
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