Castilleja chromosa |
Castilleja mendocinensis |
|
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desert paintbrush |
Mendocino coast Indian paintbrush, Mendocino coast paintbrush |
|
Habit | Herbs, sometimes subshrubs, perennial, 1.5–3.5(–4.5) dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.7–6.5 dm; from a woody caudex; with a taproot. |
Stems | several to many, ascending to erect, often grayish, unbranched, rarely branched, sometimes with short, leafy axillary branches, hairs spreading-erect, long, stiff, eglandular, sometimes also with shorter, stipitate-glandular ones. |
few to many, decumbent to ascending, much-branched, with leafy axillary shoots, villous, hairs spreading, long, stiff to soft, eglandular, mixed with short-glandular ones. |
Leaves | gray-green, linear, lanceolate, or oblanceolate, sometimes broadly lanceolate, (1.5–)2.5–6(–7) cm, not fleshy, margins plane, involute, (0–)3–5(–7)-lobed, sometimes with secondary lobes, apex acuminate to obtuse; lobes spreading, linear, apex acuminate. |
gray-green becoming ± purple, or green, ± cup-shaped, oblong to narrowly elliptic or suborbicular, 0.5–2(–5) cm, ± fleshy, cupulate throughout, sometimes obscurely so on distal portion of stem, margins plane, flat to involute, 0–3-lobed, apex rounded; lobes ascending, oblong to rounded, apex truncate or rounded, sometimes ± acute. |
Inflorescences | 2.5–15 (much longer in fruit) × 1.5–5.5 cm; bracts proximally greenish to dull purplish, distally bright red to scarlet or orange-red, rarely yellowish to dull orange or pink, narrowly to broadly linear or lanceolate, narrowly ovate, or oblong-lanceolate, (0–)3–7-lobed, rarely with secondary lobes; lobes spreading, linear to oblong, sometimes oblanceolate, often expanded near tip, long, proximal lobes arising below mid length, apex rounded or obtuse to sometimes acute. |
5–23 × 1.5–3.5 cm; bracts proximally greenish, distally bright red or red-orange, sometimes orange, oblong, ovate, or widely cuneate to widely obovate to suborbiculate, sometimes cup-shaped, 0–3-lobed, sometimes with 3 additional shallow teeth at tip of central lobe; lobes erect, oblong to broadly triangular, short, arising above mid length, apex truncate, rounded, or obtuse, lateral ones sometimes acute. |
Corollas | straight or ± curved, 18–35(–40) mm; tube 8–15 mm; beak short- or long-exserted, adaxially green to yellow-green, (9–)10–18 mm; abaxial lip deep green, reduced, thickened, included to exserted, 2–3 mm, ca. 20% as long as beak; teeth incurved, deep green, 0.5–1 mm. |
straight or slightly curved, 28–45 mm; tube 18–20 mm; beak exserted, adaxially green or yellow-green, 15–25 mm; abaxial lip deep green, reduced, visible in front cleft, 1–1.5 mm, 10% as long as beak; teeth incurved, green, 0.5–1 mm. |
Calyces | colored as bracts, sometimes with broad yellow band below colored lobe apices, (17–)20–27 mm; abaxial clefts 4–10 mm, adaxial 6–12 mm, abaxial ca. 30% of calyx length, adaxial ca. 40% of calyx length, deeper than laterals, lateral 1–4 mm, ca. 15% of calyx length; lobes oblong or ovate to narrowly triangular or lanceolate, apex obtuse to rounded. |
colored as bracts, often with a yellow central band, 20–31 mm; abaxial and adaxial clefts 8–12 mm, ca. 50% of calyx length, deeper than laterals, lateral 2–6 mm, 10–15% of calyx length; lobes oblong to broadly or narrowly triangular, apex obtuse or rounded, sometimes acute. |
2n | = 24, 48. |
= 72. |
Castilleja chromosa |
Castilleja mendocinensis |
|
Phenology | Flowering (Feb–)Mar–Aug(–Nov). | Flowering Apr–Aug. |
Habitat | Dry sagebrush slopes and flats, pinyon-juniper stands, blackbrush, open yellow pine forests. | Coastal scrub, headlands, sea bluffs, over sandstone or serpentine. |
Elevation | 500–3200 m. (1600–10500 ft.) | 0–100 m. (0–300 ft.) |
Distribution |
AZ; CA; CO; ID; NM; NV; OR; UT; WY
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CA; OR
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Discussion | Castilleja chromosa is sometimes confused with 3b. C. angustifolia var. dubia (see discussion there). Castilleja chromosa retains its distinctive morphology across its wide range and is a characteristic species of much of the southwestern United States. Where it overlaps with C. angustifolia, the two are distinguished by inflorescence color and width and by the lengths of the calyx, corolla, and corolla beak. In the broad region of their sympatry, there is little evidence of intergradation, except in a few sites in Elko County, Nevada, and in southern Wyoming. Throughout southern Idaho and northeastern Nevada the range of the two overlap with little or no intergradation. At high elevations in Montrose County, Colorado, C. chromosa has narrower leaves and a longer and silkier pubescence, especially in the inflorescence. Apparent hybrids between C. chromosa and C. flava var. rustica are known from Custer County, Idaho, and hybrids with C. linariifolia are known from Montrose County, Colorado. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja mendocinensis is a coastal plant from Mendocino County, California, northward to Curry County, Oregon. Its close relative, C. latifolia, occurs in similar habitats south of San Francisco Bay. There is one known population in Oregon, and a number of California localities are threatened by coastal development. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 595. | FNA vol. 17, p. 626. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. ewanii, C. martini subsp. ewanii, C. martini var. ewanii | C. latifolia subsp. mendocinensis |
Name authority | A. Nelson: Bull. Torrey Bot. Club 26: 245. (1899) — (as Castilleia) | (Eastwood) Pennell: Proc. Acad. Nat. Sci. Philadelphia 99: 184. (1947) |
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