Castilleja chromosa |
Castilleja martini |
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desert paintbrush |
Camp Martin paintbrush |
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Habit | Herbs, sometimes subshrubs, perennial, 1.5–3.5(–4.5) dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.5–5(–8) dm; from a woody caudex; with a taproot. | ||||
Stems | several to many, ascending to erect, often grayish, unbranched, rarely branched, sometimes with short, leafy axillary branches, hairs spreading-erect, long, stiff, eglandular, sometimes also with shorter, stipitate-glandular ones. |
several to many, erect or ascending, sometimes curved at base, unbranched or branched, sometimes with short, leafy axillary shoots, hairs spreading, short or long and distally short, stiff or soft, stipitate-glandular. |
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Leaves | gray-green, linear, lanceolate, or oblanceolate, sometimes broadly lanceolate, (1.5–)2.5–6(–7) cm, not fleshy, margins plane, involute, (0–)3–5(–7)-lobed, sometimes with secondary lobes, apex acuminate to obtuse; lobes spreading, linear, apex acuminate. |
green to sometimes purple, narrowly oblong or lanceolate to sometimes linear or narrowly ovate, (0.8–)1–5(–6.5) cm, not fleshy, margins wavy, flat or involute, 0–5-lobed, apex rounded or obtuse, rarely acute; lobes ascending-spreading or divaricate, linear to narrowly oblong, apex acute. |
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Inflorescences | 2.5–15 (much longer in fruit) × 1.5–5.5 cm; bracts proximally greenish to dull purplish, distally bright red to scarlet or orange-red, rarely yellowish to dull orange or pink, narrowly to broadly linear or lanceolate, narrowly ovate, or oblong-lanceolate, (0–)3–7-lobed, rarely with secondary lobes; lobes spreading, linear to oblong, sometimes oblanceolate, often expanded near tip, long, proximal lobes arising below mid length, apex rounded or obtuse to sometimes acute. |
2.5–25 × 1.5–4 cm; bracts proximally greenish, distally red, orange-red, or pale orange, rarely yellow, lanceolate or broadly lanceolate to oblong, (0–)3–5(–7)-lobed, plus 1–3 shallow teeth on apex of central lobe (var. clokeyi); lobes ascending or spreading, linear, lanceolate, or oblanceolate, medium length or long, arising at or near mid length, central lobe apex obtuse to rounded, lateral ones acute. |
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Corollas | straight or ± curved, 18–35(–40) mm; tube 8–15 mm; beak short- or long-exserted, adaxially green to yellow-green, (9–)10–18 mm; abaxial lip deep green, reduced, thickened, included to exserted, 2–3 mm, ca. 20% as long as beak; teeth incurved, deep green, 0.5–1 mm. |
straight to slightly curved, (19–)22–45 mm; tube 11–22 mm; abaxial lip rarely exserted, beak exserted; beak adaxially green to yellowish, 9–21(–23) mm; abaxial lip green or deep green with white teeth or red with green teeth, reduced, not inflated, often visible through front cleft, 1–2 mm, ca. 10% as long as beak; teeth incurved, green to white, 0.5–1 mm. |
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Calyces | colored as bracts, sometimes with broad yellow band below colored lobe apices, (17–)20–27 mm; abaxial clefts 4–10 mm, adaxial 6–12 mm, abaxial ca. 30% of calyx length, adaxial ca. 40% of calyx length, deeper than laterals, lateral 1–4 mm, ca. 15% of calyx length; lobes oblong or ovate to narrowly triangular or lanceolate, apex obtuse to rounded. |
proximally pale green, green, or whitish, sometimes with a yellow central band, distally colored as bracts, 14–26(–28) mm; abaxial and adaxial clefts 3–10 mm, 20–50% of calyx length, deeper than laterals, lateral 1–5 mm, 15–20% of calyx length; lobes lanceolate to broadly lanceolate, oblong, or ovate, apex acute, obtuse, or rounded. |
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2n | = 24, 48. |
= 24, 48, 72. |
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Castilleja chromosa |
Castilleja martini |
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Phenology | Flowering (Feb–)Mar–Aug(–Nov). | |||||
Habitat | Dry sagebrush slopes and flats, pinyon-juniper stands, blackbrush, open yellow pine forests. | |||||
Elevation | 500–3200 m. (1600–10500 ft.) | |||||
Distribution |
AZ; CA; CO; ID; NM; NV; OR; UT; WY
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CA; NV; nw Mexico |
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Discussion | Castilleja chromosa is sometimes confused with 3b. C. angustifolia var. dubia (see discussion there). Castilleja chromosa retains its distinctive morphology across its wide range and is a characteristic species of much of the southwestern United States. Where it overlaps with C. angustifolia, the two are distinguished by inflorescence color and width and by the lengths of the calyx, corolla, and corolla beak. In the broad region of their sympatry, there is little evidence of intergradation, except in a few sites in Elko County, Nevada, and in southern Wyoming. Throughout southern Idaho and northeastern Nevada the range of the two overlap with little or no intergradation. At high elevations in Montrose County, Colorado, C. chromosa has narrower leaves and a longer and silkier pubescence, especially in the inflorescence. Apparent hybrids between C. chromosa and C. flava var. rustica are known from Custer County, Idaho, and hybrids with C. linariifolia are known from Montrose County, Colorado. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja martini is sometimes treated as a subspecies of C. applegatei, but there are relatively consistent morphological discontinuities between these two variable complexes. See notes under 4. C. applegatei. Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 595. | FNA vol. 17, p. 625. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | C. ewanii, C. martini subsp. ewanii, C. martini var. ewanii | C. applegatei subsp. martini | ||||
Name authority | A. Nelson: Bull. Torrey Bot. Club 26: 245. (1899) — (as Castilleia) | Abrams: Bull. S. Calif. Acad. Sci. 1: 69. (1902) | ||||
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