Castilleja chambersii |
Castilleja uliginosa |
|
---|---|---|
Chambers' Indian paintbrush, Chambers' paintbrush |
Pitkin Marsh Indian paintbrush, Pitkin Marsh paintbrush |
|
Habit | Herbs, perennial, 1.5–3.7 dm; from a thick, woody caudex; with a taproot. | Herbs, perennial, 3–5 dm; from a woody caudex; rhizomatous. |
Stems | solitary or few to several, ascending, sometimes short-decumbent and rooting, branched from near base, sometimes distally, glabrous or glabrate with hairs very sparse, ± appressed, very short, soft, eglandular. |
several, decumbent proximally and sometimes becoming weakly rhizomatous, becoming ascending to erect, unbranched or often branched proximally, hairs spreading, short and long, soft, mixed eglandular and stipitate-glandular. |
Leaves | green, often brown- or purple-tinged, oblong to lanceolate to narrowly ovate or ovate, (1–)2–5.8 cm, not fleshy, margins plane, ± involute, deeply 3–7(–11)-lobed, sometimes with secondary lobes, apex narrowly acute; lobes erect to ascending, lanceolate to linear-lanceolate to oblanceolate, apex acute. |
green, lanceolate or broadly lanceolate, 3–5 cm, much reduced distally, not fleshy, margins plane, sometimes ± wavy, flat, 0-lobed, apex acute to rounded. |
Inflorescences | 3–15 × 2–4.5 cm; bracts proximally greenish, distally bright red, scarlet, or pale reddish orange, rarely orange-yellow, often fading to pale yellowish orange with age, obovate to orbicular, fan-shaped, (3–)5–9(–13)-lobed, sometimes with secondary lobes; lobes erect or ascending, lanceolate to triangular, short and medium length, usually arising at or above mid length, rarely below, apex acute. |
10–30 × 3–4 cm; bracts pale yellow to cream throughout, or proximally pale greenish near base, distally pale yellow to cream, broadly lanceolate, 3-lobed; lobes ascending, narrowly lanceolate, long, arising near mid length, apex narrowly acute or acuminate. |
Corollas | straight, 30–45 mm; tube 14–19 mm; beak long-exserted, adaxially green or yellow-green to brownish, 18–24 mm; abaxial lip deep green, reduced, 1–3 mm, 10% as long as beak; teeth incurved, greenish to dull purplish, 0.5–1.5 mm. |
slightly curved, 22–30 mm; tube 11–15 mm; beak partly exserted, adaxially whitish, yellowish, or greenish, 10–15 mm; abaxial lip green, small, protuberant, 3 mm, ca. 20% as long as beak; teeth erect to incurved, green, to 1 mm. |
Calyces | proximally green, sometimes purple to brown, distally colored as bracts, 20–30 mm; abaxial and adaxial clefts 7–14 mm, ca. 33% of calyx length, deeper than laterals, lateral 2–4 mm, 10–15% of calyx length; lobes triangular, barely longer than wide, apex acute or acuminate to obtuse. |
colored as bracts, 20–25 mm; abaxial and adaxial clefts 7–14 mm, ca. 50% of calyx length, deeper than laterals, lateral 5–6 mm, ca. 33% of calyx length; lobes linear, apex acute, ciliate. |
2n | = 24. |
|
Castilleja chambersii |
Castilleja uliginosa |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jun. |
Habitat | Turf or crevices on rocky slopes and benches over basalt, se to sw aspect, sun or partial shade. | Margins of wet meadows, marshes, and wet thickets. |
Elevation | 600–1000 m. (2000–3300 ft.) | 40–60 m. (100–200 ft.) |
Distribution |
OR; WA
|
CA |
Discussion | Castilleja chambersii is limited to the summits of three volcanic peaks in the northern Coast Range of Clatsop County, Oregon, and at one similar area in nearby Pacific County, Washington. It is similar to C. rupicola, and the two likely share a common ancestor. Disturbance and erosion from logging and road construction represent significant threats to C. chambersii. Populations of C. chambersii often grow near and even alongside C. hispida, but hybrids are rare. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja uliginosa is endemic to Pitkin Marsh in Sonoma County. It differs from C. miniata by its uniformly pale yellow inflorescences and pubescent stems, as well as by its disjunct, low-elevation habitat. Much of its available habitat was destroyed by development, and it is apparently now extirpated from the wild. It survives in the form of tissue clones from the last wild plant, backcrossed with C. miniata by L. R. Heckard in the 1980s and maintained since then in the greenhouses at the University of California at Berkeley. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 593. | FNA vol. 17, p. 662. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Name authority | J. M. Egger & Meinke: Brittonia 51: 445, fig. 1. (1999) | Eastwood: Leafl. W. Bot. 3: 117. (1942) |
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