Castilleja chambersii |
Castilleja campestris |
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Chambers' Indian paintbrush, Chambers' paintbrush |
field owl's-clover, meadow paintbrush, vernal pool Indian paintbrush, yellow owl's clover |
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Habit | Herbs, perennial, 1.5–3.7 dm; from a thick, woody caudex; with a taproot. | Herbs, annual, (0.3–)0.9–3 dm; with fibrous roots. | ||||
Stems | solitary or few to several, ascending, sometimes short-decumbent and rooting, branched from near base, sometimes distally, glabrous or glabrate with hairs very sparse, ± appressed, very short, soft, eglandular. |
solitary or few, erect, unbranched, sometimes branched, glabrous or hairs sparse proximally, spreading, long, soft distally. |
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Leaves | green, often brown- or purple-tinged, oblong to lanceolate to narrowly ovate or ovate, (1–)2–5.8 cm, not fleshy, margins plane, ± involute, deeply 3–7(–11)-lobed, sometimes with secondary lobes, apex narrowly acute; lobes erect to ascending, lanceolate to linear-lanceolate to oblanceolate, apex acute. |
green, sometimes purple-tinged, linear to narrowly lanceolate, (0.4–)1–4 cm, not fleshy, thin and flexible or thick and ± brittle, margins plane, involute, 0-lobed, apex acuminate. |
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Inflorescences | 3–15 × 2–4.5 cm; bracts proximally greenish, distally bright red, scarlet, or pale reddish orange, rarely orange-yellow, often fading to pale yellowish orange with age, obovate to orbicular, fan-shaped, (3–)5–9(–13)-lobed, sometimes with secondary lobes; lobes erect or ascending, lanceolate to triangular, short and medium length, usually arising at or above mid length, rarely below, apex acute. |
2–15 × 2–3 cm; bracts green throughout, sometimes purplish tinged, linear to narrowly lanceolate, 0-lobed, apex acute. |
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Corollas | straight, 30–45 mm; tube 14–19 mm; beak long-exserted, adaxially green or yellow-green to brownish, 18–24 mm; abaxial lip deep green, reduced, 1–3 mm, 10% as long as beak; teeth incurved, greenish to dull purplish, 0.5–1.5 mm. |
straight, 11–26 mm; tube 10–22 mm; abaxial lip, beak, and part of tube exserted from calyx; beak adaxially white or yellow, 3–6 mm; abaxial lip yellow, sometimes orange, paler near teeth, prominent, deeply saccate with 3 lobes at ca. 90° from one another, pouches 6–10 mm wide, 3–4 mm deep, abruptly widening from tube, sacs ± round or ± rounded-triangular from above, villous within at base of teeth, 4–5 mm (2–4 mm from sinus), 70–90% as long as beak, puberulent; teeth erect, white, sometimes with deeply colored bases, 0.5–2 mm. |
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Calyces | proximally green, sometimes purple to brown, distally colored as bracts, 20–30 mm; abaxial and adaxial clefts 7–14 mm, ca. 33% of calyx length, deeper than laterals, lateral 2–4 mm, 10–15% of calyx length; lobes triangular, barely longer than wide, apex acute or acuminate to obtuse. |
proximally green, pale, or purplish, distally purple or green, 5.5–11 mm; abaxial and adaxial clefts 2–5 mm, 30–50% of calyx length, deeper than laterals, lateral 1.5–2.5 mm, 25–30% of calyx length; lobes linear-lanceolate to narrowly triangular, apex acute to acuminate. |
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2n | = 24. |
= 24. |
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Castilleja chambersii |
Castilleja campestris |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Turf or crevices on rocky slopes and benches over basalt, se to sw aspect, sun or partial shade. | |||||
Elevation | 600–1000 m. (2000–3300 ft.) | |||||
Distribution |
OR; WA
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CA; OR
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Discussion | Castilleja chambersii is limited to the summits of three volcanic peaks in the northern Coast Range of Clatsop County, Oregon, and at one similar area in nearby Pacific County, Washington. It is similar to C. rupicola, and the two likely share a common ancestor. Disturbance and erosion from logging and road construction represent significant threats to C. chambersii. Populations of C. chambersii often grow near and even alongside C. hispida, but hybrids are rare. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Castilleja campestris is sometimes confused with Triphysaria eriantha subsp. eriantha, but C. campestris has two-celled anthers and entire leaves, while T. eriantha has one pollen sac per stamen and strongly divided leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 593. | FNA vol. 17, p. 592. | ||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Orthocarpus campestris | |||||
Name authority | J. M. Egger & Meinke: Brittonia 51: 445, fig. 1. (1999) | (Bentham) T. I. Chuang & Heckard: Syst. Bot. 16: 656. (1991) | ||||
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