Castilleja brevilobata |
Castilleja fraterna |
|
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short-lobed paintbrush |
fraternal Indian paintbrush, fraternal paintbrush |
|
Habit | Herbs, perennial, 1–5 dm; from a branched, woody caudex; with a taproot. | Herbs, perennial, 0.8–2.5 dm; from a woody caudex; with a taproot. |
Stems | solitary or few, erect or ascending, unbranched, sometimes branched, hairs spreading, short, medium, and long, soft, short and medium ones short stipitate-glandular. |
few to many, ± curved at base, ascending or erect distally, unbranched, hairs moderately dense, spreading, medium length and long, soft, mixed with shorter stipitate-glandular ones. |
Leaves | green or ± yellow, lanceolate, elliptic, or oblong to narrowly ovate, 1–2(–2.5) cm, not fleshy, margins plane or wavy, involute, (0–)3–5(–7)-lobed, apex rounded to acute; lobes ascending to erect, linear to lanceolate, apex rounded to acute. |
green, sometimes purple-tinged, lanceolate, broadly lanceolate, or narrowly ovate, 1–4.3 cm, not fleshy, margins plane, sometimes ± wavy, ± involute, 0–3-lobed, apex acute to acuminate; lobes ascending, often narrowly oblanceolate, apex acute to acuminate. |
Inflorescences | 3–20 × 2–3.5 cm; bracts proximally greenish to dull brown, distally red, orange-red, or scarlet, sometimes orange or yellow, broadly lanceolate or oblong, (0–)3–5-lobed; lobes ascending, broadly to narrowly lanceolate, short, arising above mid length, apex acute, obtuse, or rounded. |
3–8.5 × 1–4 cm; bracts greenish or bright to sometimes dull red, sometimes scarlet, orange, or pale pink throughout, or proximally greenish, distally as above, ovate to elliptic, (0–)3-lobed; lobes ascending, lanceolate to narrowly triangular, short, arising above mid length, apex acute to obtuse or acuminate. |
Corollas | straight, 15–24(–26) mm; tube 12–16 mm; beak exserted, abaxial lip equal to calyx; beak adaxially green or ± yellow-green, 7–10 mm, puberulent, stipitate-glandular; abaxial lip deep green, reduced, rounded, 1–2 mm, 10–25% as long as beak; teeth incurved to erect, light green, 0.5–1 mm. |
straight or slightly curved, 20–40 mm; tube 27 mm; abaxial lip often visible in abaxial cleft, sometimes exserted, beak usually exserted from calyx; beak adaxially green to yellow, 8–14 mm; abaxial lip red, black, green, white, or green and white, ± prominent, slightly rounded, 1.5–5 mm, ca. 33% as long as beak; teeth ascending, green, bright red, pink, white, or yellow, 0.5–3 mm. |
Calyces | green or whitish with green veins, lobes colored as bract lobes or paler, 14–30 mm; abaxial and adaxial clefts 5.5–8.5 mm, 30–40% of calyx length, deeper than laterals, lateral 1.5–4 mm, 20–25% of calyx length; lobes oblong to narrowly triangular, apex obtuse to rounded. |
brightly, conspicuously colored for at least distal 2/3, often throughout, providing much of inflorescence coloration, proximally pale whitish to pale pink, distally colored as in distal portion of bracts, sometimes colored throughout as in distal portion of bracts, 15–30 mm; abaxial and adaxial clefts 7–14 mm, 33–50% of calyx length, deeper than laterals, lateral 1–5 mm, 5–15% of calyx length; lobes narrowly to broadly triangular, apex rounded to acute. |
2n | = 24. |
|
Castilleja brevilobata |
Castilleja fraterna |
|
Phenology | Flowering Mar–Aug. | Flowering Jul–Aug(–Sep). |
Habitat | Dry savannas, rocky slopes and open conifer forests, on serpentine. | Moist or dry rocky slopes and flats, ridges, talus, dwarf willow mats, subalpine to alpine, rarely along stream channels at lower elevations, over sedimentary rocks, often limy. |
Elevation | 200–1700 m. (700–5600 ft.) | 2000–2900 m. (6600–9500 ft.) |
Distribution |
CA; OR
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OR
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Discussion | Castilleja brevilobata is endemic to dry serpentine openings in the Siskiyou Mountains of southwestern Oregon and adjacent California. Although sometimes treated as part of C. applegatei or C. hispida, its morphology does not suggest a close connection with either. This species occasionally hybridizes with C. pruinosa in Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja fraterna is endemic to one ridge system in the Wallowa Mountains of northeastern Oregon. A parallel ridge system in the range has a second Wallowa Mountains endemic, C. rubida. Castilleja fraterna is colored similarly to C. miniata, but the petaloid teeth of the abaxial corolla lip are more like C. chrysantha. It is possible that C. fraterna was derived through hybridization, though its chromosome number is unknown. It resembles the newly described C. kerryana in Montana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 591. | FNA vol. 17, p. 609. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. hispida subsp. brevilobata | |
Name authority | Piper: Proc. Biol. Soc. Wash. 33: 104. (1920) | Greenman: Bot. Gaz. 48: 147. (1909) |
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