Castilleja brevilobata |
Castilleja applegatei |
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short-lobed paintbrush |
Applegate's Indian paintbrush, Applegate's paintbrush, wavy leaf paintbrush, wavy-leaf Indian paintbrush, waxy-leaf Indian paintbrush |
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Habit | Herbs, perennial, 1–5 dm; from a branched, woody caudex; with a taproot. | Herbs, perennial, (0.8–)1–5(–6) dm; from a branched, woody caudex; with a taproot. | ||||||||||||
Stems | solitary or few, erect or ascending, unbranched, sometimes branched, hairs spreading, short, medium, and long, soft, short and medium ones short stipitate-glandular. |
few to many, erect, ascending, or decumbent, unbranched or branched, hairs sparse to dense, spreading, long, soft to ± stiff, eglandular, mixed with shorter stipitate-glandular ones. |
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Leaves | green or ± yellow, lanceolate, elliptic, or oblong to narrowly ovate, 1–2(–2.5) cm, not fleshy, margins plane or wavy, involute, (0–)3–5(–7)-lobed, apex rounded to acute; lobes ascending to erect, linear to lanceolate, apex rounded to acute. |
green to purplish or brown, linear to broadly lanceolate, sometimes ovate, 1–6 cm, not fleshy, margins wavy, involute, (0–)3(–5)-lobed, apex rounded or narrowly acute to acuminate; lobes spreading or ascending, lanceolate to broadly lanceolate, often short, apex acute to rounded. |
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Inflorescences | 3–20 × 2–3.5 cm; bracts proximally greenish to dull brown, distally red, orange-red, or scarlet, sometimes orange or yellow, broadly lanceolate or oblong, (0–)3–5-lobed; lobes ascending, broadly to narrowly lanceolate, short, arising above mid length, apex acute, obtuse, or rounded. |
2–12(–21 in fruit) × 1–5 cm; bracts proximally green to dull purplish brown, distally red, red-orange, or scarlet, sometimes orange, white, or yellow, rarely with a narrow yellowish band medially, lanceolate, broadly lanceolate, oblong, or lanceolate-ovate, 3–5(–7)-lobed; lobes spreading to ascending, linear, sometimes expanded near tip, long, arising from ca. mid length, central lobe apex obtuse to rounded, sometimes expanded, others acute, rounded, obtuse, or acuminate. |
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Corollas | straight, 15–24(–26) mm; tube 12–16 mm; beak exserted, abaxial lip equal to calyx; beak adaxially green or ± yellow-green, 7–10 mm, puberulent, stipitate-glandular; abaxial lip deep green, reduced, rounded, 1–2 mm, 10–25% as long as beak; teeth incurved to erect, light green, 0.5–1 mm. |
straight or curved in proximal 1/3, 16–35(–41) mm; tube 9–22 mm; beak usually long-exserted, adaxially green, yellow-green, or yellow, 6–20 mm; abaxial lip deep green to whitish or yellow, reduced, inconspicuous, protuberant, thickened, included or exserted, 1–3 mm, 5–20% as long as beak; teeth ascending or incurved, deep green to yellow, 0.5–1 mm. |
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Calyces | green or whitish with green veins, lobes colored as bract lobes or paler, 14–30 mm; abaxial and adaxial clefts 5.5–8.5 mm, 30–40% of calyx length, deeper than laterals, lateral 1.5–4 mm, 20–25% of calyx length; lobes oblong to narrowly triangular, apex obtuse to rounded. |
proximally green or whitish, sometimes yellow, lobes colored as bract lobes, sometimes with a yellow band between proximal and distal portions, 13–25 mm; abaxial and adaxial clefts 4–14 mm, 33–50% of calyx length, deeper than laterals, lateral 2.5–8 mm, 12–50% of calyx length; lobes lanceolate-acuminate, narrowly oblong, or narrowly triangular, apex acuminate, acute, or obtuse. |
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2n | = 24. |
= 24, 48. |
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Castilleja brevilobata |
Castilleja applegatei |
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Phenology | Flowering Mar–Aug. | |||||||||||||
Habitat | Dry savannas, rocky slopes and open conifer forests, on serpentine. | |||||||||||||
Elevation | 200–1700 m. (700–5600 ft.) | |||||||||||||
Distribution |
CA; OR
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CA; ID; NV; OR; UT; WY
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Discussion | Castilleja brevilobata is endemic to dry serpentine openings in the Siskiyou Mountains of southwestern Oregon and adjacent California. Although sometimes treated as part of C. applegatei or C. hispida, its morphology does not suggest a close connection with either. This species occasionally hybridizes with C. pruinosa in Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Castilleja applegatei is a widespread and often common species, with complex patterns of variation and several common but inconstant color forms, especially in var. pinetorum. Castilleja disticha and C. martini, although sometimes included as subspecies within C. applegatei, are treated as species here. Both are more morphologically divergent and more easily distinguished from typical C. applegatei than are the four varieties accepted here. Variety pinetorum is the most widespread form and also occurs over a wider range of elevations than the other three varieties, which are primarily montane to subalpine and occasionally alpine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 591. | FNA vol. 17, p. 587. | ||||||||||||
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja | ||||||||||||
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Synonyms | C. hispida subsp. brevilobata | |||||||||||||
Name authority | Piper: Proc. Biol. Soc. Wash. 33: 104. (1920) | Fernald: Erythea 6: 49. (1898) — (as Castilleia) | ||||||||||||
Web links |