Castilleja attenuata |
Castilleja peckiana |
|
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attenuate Indian paintbrush, attenuate paintbrush, narrow-leaf owl's-clover, narrow-leaf owl-clover, narrow-leaf paintbrush, valley-tassels |
peak paintbrush, Peck's Indian paintbrush, Peck's paintbrush |
|
Habit | Herbs, annual, 1–4.5 dm; with fibrous roots. | Herbs, perennial, (1.8–)2.4–6 dm; from a woody caudex; with a stout taproot. |
Stems | solitary, erect to ascending, unbranched, sometimes branched from base, hairs spreading, short and medium, ± stiff, eglandular. |
few to many, erect or ascending, often branched distally, sometimes unbranched, sometimes with short axillary shoots, proximal hairs retrorse to appressed, short to moderately long, distal hairs spreading, longer, soft, often mixed with short stipitate-glandular ones. |
Leaves | green to purple-tinged, linear to linear-lanceolate, (1–)2–8 cm, not fleshy, margins plane, flat, 0(–5)-lobed, apex acuminate; lobes spreading to ascending, filiform to lanceolate, apex sometimes acuminate. |
green, linear-lanceolate, rarely broadly lanceolate, (1.2–)2.5–8(–9) cm, not fleshy, margins plane, flat or involute, 0(–3)-lobed, apex acute to rounded; lobes ascending-spreading, narrowly lanceolate to linear, apex acute or obtuse. |
Inflorescences | (1.5–)2–10(–19, –30 in fruit) × 1–2 cm; bracts proximally green to pale brown, rarely dull reddish brown, distally white on apices, sometimes pale yellow or pale pink-purplish on apices, rarely greenish or dull reddish brown throughout, lanceolate or lanceolate-elliptic, 3(–5)-lobed; lobes ascending, linear to lanceolate, long, arising below mid length, apex acuminate, acute, or obtuse. |
(2–)4–17 × 1.5–3 cm; bracts proximally greenish, distally red, orange-red, or orange, proximal sometimes lanceolate, distal broadly lanceolate to ovate, (0–)3(–7)-lobed; lobes spreading to ascending, linear to lanceolate, long, arising near or below mid length, central lobe apex rounded, lateral ones acute. |
Corollas | straight, 10–25 mm; tube 9–20 mm, not expanded distally; beak exserted, straight, adaxially white, light yellow, or greenish, 3–5 mm, inconspicuously puberulent; abaxial lip white or yellow with deep brown to purple spots, often becoming pink, slightly inflated, exserted or not, pouches 3, 2 mm wide, 1–1.5 mm deep, 3–4 mm, 75–80% as long as beak; teeth erect, white, pale yellow, or pink, 0.5–1.2 mm. |
straight, 23–30(–35) mm; tube 12–20 mm; beak exserted, adaxially green, 8–12(–14) mm; abaxial lip pale to deep green, reduced, rounded, 0.5–1.5 mm, 10–20% as long as beak; teeth erect to incurved, green, (0.4–)0.7–1.2(–2) mm. |
Calyces | colored as bracts, (8–)15–23 mm; abaxial and adaxial clefts (4–)6–8 mm, abaxial ca. 50% of calyx length, adaxial ca. 75% of calyx length, lateral 3–3.5 mm, 33% of calyx length; lobes linear to narrowly triangular, apex acute to acuminate. |
proximally pale yellow or greenish, distally colored as bracts, (15–)18–28 mm; abaxial and adaxial clefts 6–12 mm, 40–45% of calyx length, deeper than laterals, lateral 2.5–8 mm, 20–30% of calyx length; lobes linear to narrowly lanceolate, apex acute to acuminate. |
Filaments | glabrous. |
|
2n | = 24. |
= 72, 96, ca. 120. |
Castilleja attenuata |
Castilleja peckiana |
|
Phenology | Flowering (Feb–)Mar–Jun. | Flowering (Apr–)May–Aug. |
Habitat | Grasslands, pastures, moist margins of springs and streams, damp rocky slopes. | Open conifer forests, sagebrush slopes, riparian meadows, shores. |
Elevation | 0–2100 m. (0–6900 ft.) | 1400–2600 m. (4600–8500 ft.) |
Distribution |
AZ; CA; OR; WA; BC; Mexico (Baja California); South America (Chile)
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ID; NV; OR
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Discussion | Castilleja attenuata is a common and widespread species, ranging from southwestern Canada to northern Baja California, with several disjunct populations in central Chile. It is sensitive to competition from weeds. Disjunct populations in the Rincon Mountains in Pima County, Arizona, often have pink bracts but are otherwise typical. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
When describing Castilleja peckiana, Pennell noted that its variation approached C. hispida on one hand and C. miniata on the other, and it is likely of allopolyploid derivation. L. R. Heckard (1968) found chromosome numbers of n = 36, 48, and ca. 60. He hypothesized that C. chromosa, C. hispida var. acuta, and C. miniata were likely involved in its ancestry, and possibly C. pruinosa as well. Heckard suggested subsequent introgression among the derived forms introduced further complexity. Though complex, these forms are self-perpetuating and appear morphologically stable within their range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 590. | FNA vol. 17, p. 643. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | Orthocarpus attenuatus | |
Name authority | (A. Gray) T. I. Chuang & Heckard: Syst. Bot. 16: 656. (1991) | Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 9. (1941) |
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