Castilleja attenuata |
Castilleja lanata |
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attenuate Indian paintbrush, attenuate paintbrush, narrow-leaf owl's-clover, narrow-leaf owl-clover, narrow-leaf paintbrush, valley-tassels |
Sierra woolly Indian paintbrush, Sierra woolly paintbrush, woolly paintbrush |
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Habit | Herbs, annual, 1–4.5 dm; with fibrous roots. | Subshrubs or herbs, perennial, 1.8–9(–10) dm; from a woody caudex; with a thick, woody taproot or system to thick, branched roots. |
Stems | solitary, erect to ascending, unbranched, sometimes branched from base, hairs spreading, short and medium, ± stiff, eglandular. |
few to many, erect to ascending, unbranched, sometimes branched distally, sometimes with leafy axillary shoots, lanate to woolly, hairs dense, spreading to weakly appressed, white to yellowish, medium length, soft, moderately branched or unbranched in different populations, sometimes mixed, eglandular, rarely short stipitate-glandular, mostly obscuring surface. |
Leaves | green to purple-tinged, linear to linear-lanceolate, (1–)2–8 cm, not fleshy, margins plane, flat, 0(–5)-lobed, apex acuminate; lobes spreading to ascending, filiform to lanceolate, apex sometimes acuminate. |
green to purple, often hidden by matted, white hairs, linear to narrowly oblong or narrowly lanceolate, (1–)2–7(–10) cm, not fleshy, margins plane, flat, sometimes involute, 0–3-lobed, apex rounded, sometimes acute; lobes spreading, linear or narrowly oblong, apex rounded to acute. |
Inflorescences | (1.5–)2–10(–19, –30 in fruit) × 1–2 cm; bracts proximally green to pale brown, rarely dull reddish brown, distally white on apices, sometimes pale yellow or pale pink-purplish on apices, rarely greenish or dull reddish brown throughout, lanceolate or lanceolate-elliptic, 3(–5)-lobed; lobes ascending, linear to lanceolate, long, arising below mid length, apex acuminate, acute, or obtuse. |
2–19 × 1.5–5 cm; bracts proximally pale greenish to greenish gray, distally bright red to orange-red, sometimes pinkish, magenta, or salmon, rarely yellow, oblong to narrowly lanceolate or narrowly oblanceolate, deeply 0–3(–5)-lobed; lobes spreading, oblanceolate or linear, long, arising below mid length, distal bract apex usually rounded. |
Corollas | straight, 10–25 mm; tube 9–20 mm, not expanded distally; beak exserted, straight, adaxially white, light yellow, or greenish, 3–5 mm, inconspicuously puberulent; abaxial lip white or yellow with deep brown to purple spots, often becoming pink, slightly inflated, exserted or not, pouches 3, 2 mm wide, 1–1.5 mm deep, 3–4 mm, 75–80% as long as beak; teeth erect, white, pale yellow, or pink, 0.5–1.2 mm. |
curved in proximal 1/3, 23–35(–42) mm; tube 12–17 mm; beak exserted, adaxially greenish to yellowish, 11–22 mm; abaxial lip dark green, reduced, 1–4 mm, ca. 20% as long as beak; teeth incurved, reduced, green, 0.5–2.5 mm. |
Calyces | colored as bracts, (8–)15–23 mm; abaxial and adaxial clefts (4–)6–8 mm, abaxial ca. 50% of calyx length, adaxial ca. 75% of calyx length, lateral 3–3.5 mm, 33% of calyx length; lobes linear to narrowly triangular, apex acute to acuminate. |
proximally whitish to green, central band whitish, distal 1/3 colored as bract lobes, sometimes lighter or deeper, 15–29 mm; abaxial and adaxial clefts 6–14 mm, ca. 50% of calyx length, deeper than laterals, lateral 0–5 mm, 0–5(–15)% of calyx length; lobes oblong to broadly triangular, apex rounded, truncate, or emarginate. |
Filaments | glabrous. |
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2n | = 24. |
= 24. |
Castilleja attenuata |
Castilleja lanata |
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Phenology | Flowering (Feb–)Mar–Jun. | Flowering year-round. |
Habitat | Grasslands, pastures, moist margins of springs and streams, damp rocky slopes. | Dry rocky slopes, ridges, canyons, flats, valleys, montane. |
Elevation | 0–2100 m. (0–6900 ft.) | 300–2300 m. (1000–7500 ft.) |
Distribution |
AZ; CA; OR; WA; BC; Mexico (Baja California); South America (Chile)
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AZ; NM; TX; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
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Discussion | Castilleja attenuata is a common and widespread species, ranging from southwestern Canada to northern Baja California, with several disjunct populations in central Chile. It is sensitive to competition from weeds. Disjunct populations in the Rincon Mountains in Pima County, Arizona, often have pink bracts but are otherwise typical. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most records of Castilleja lanata are from the southern third of Arizona, New Mexico, and southwestern Texas. Plants sometimes referred to C. lanata from the Channel Islands of California are assigned here to C. hololeuca, an insular endemic. In northern Mexico, C. lanata extends from San Luis Potosí and Nuevo León westward to Sonora, with a disjunct distribution in central Baja California, where it overlaps the southern limits of a related species, C. foliolosa. Castilleja lanata inhabits a wide spectrum of soils and elevations and does not co-occur with the closely related C. galehintoniae in Nuevo León, where the latter is limited to limestone and gypsum substrates in a narrow elevational range. Hybrids of C. lanata with C. sessiliflora are known from Pecos County, Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 590. | FNA vol. 17, p. 618. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | Orthocarpus attenuatus | |
Name authority | (A. Gray) T. I. Chuang & Heckard: Syst. Bot. 16: 656. (1991) | A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 118. (1859) |
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